Rhytidaspis nigropunctata, Ingrisch, 2019

Ingrisch, Sigfrid, 2019, Revision of the genus Rhytidaspis Redtenbacher, 1891 including the description of a new genus Haudrhytidaspis gen. nov. (Orthoptera: Tettigoniidae: Conocephalinae), Zootaxa 4661 (2), pp. 343-370 : 367

Rhytidaspis nigropunctata sp. n.

Figs. 1 F–G, 2O, 3O–P, 4G, 5C, 6H, 7T–U, 8G, 9G

Holotype (male): Indonesia, Papua Barat, Raja Ampat Islands: Waigeo Island [Waigen], Mt. Nok , [0°6‘S, 130°45‘E], 1–30.vi.1938, leg. L.E. Cheesman—(London, BMNH).

Diagnosis. R. nigropunctata sp. n. is striking for its coloration: the pale face blotched with scattered black dots is unique within the genus. The stridulatory file on the underside of the left tegmen is with 2.4 mm about as short as in R. variata brevis ssp. n., but the density of the stridulatory teeth on that file with 38 teeth per mm in the middle of the file is the highest within the genus. The titillators of R. nigropunctata have an extra projection between the granular extension above the U-shaped curvature and the tip of the titillators that is strongly widened with little lobular margin not narrow as in R. picta and R. arfak sp. n. The shape of the male cerci with a sub-truncate dorsal apical lobe that has only the proximal angle triangularly projecting, and a prolonged and dorsally swollen ventral apical lobe resemble the situation in R. camela sp. n.; R. nigropunctata differs by the projecting proximal angle of the dorsal apical lobe that is less projecting but down-curved on proximal side while it is larger and flat in R. camela .

Description male. Fastigium verticis up-curved and laterally compressed, with tubercle on dorsal side in subbasal area, in apical area narrow and faintly downcurved, tip acute; on ventral side separated from fastigium frontis by a step; length of fastigium verticis before eyes 3.0 mm, greatest eye diameter 1.9 mm. Tegmen in apical half triangular with rounded tip ( Figs. 3 O–P). Stridulatory file on underside of left tegmen straight in middle, curved at both sides; 2.4 mm long, with about 105 stridulatory teeth, in middle of file with 38 teeth per mm ( Fig. 4G ). Mirror of right tegmen about oval with both lateral tips sub-angular, about 1.5 mm long, 1.9 mm wide.

Cerci with dorsal apical lobe curved mediad, terminating into a broad, sub-truncate but little irregular apical margin that has the proximal angle little but distinctly projecting and the distal angle very faintly produced ( Fig. 5C ); the ventral apical lobe is slightly narrowed in basal area behind the transverse bulge ( Fig. 7U ), afterwards with faintly concave ventral surface and distinctly convex dorsal surface; apical margin faintly sinuate with a small curved tooth from about middle of margin pointing mediad ( Fig. 6H ). Titillators with basal area curved; above the U-shaped curvature in mid-length with a granular extension that has the upper margin little lobular and is roundly extended on proximal side, while on distal side it is extended and almost reaches to tip of titillators but is subdivided by a V-shaped incision before apical third; the apical branch of the titillators is covered over the whole length with strong teeth; the extra apical sclerites are curved but simple, the apical lateral sclerites roughly oval ( Figs. 8G , 9G ).

Coloration. The dried specimen is rather uniformly light yellowish brown; pronotum, labrum and mandibles are orange brown ( Fig. 1 F–G). Face light yellow with impressed dots black Fig. 2O ); meso- and metapleura darkened; abdominal tergites with black lateral bands; abdominal sternites black in about basal half, brown in about apical half; male subgenital plate black except for yellow apical rim and base of styli; tegmina with black cells and light veins and veinlets; hind femora with very faint dark striation in basal half.

Measurements (1 male).—Body: 40; pronotum: 12.5; tegmen: 7; hind femur: 26; anterior femur: 14 mm.

Etymology. The name of the new species refers to the black dotted face; from Latin punctata dotted.

Figures

FIGURE 1. A–B, Rhytidaspis picta Redt., 1891 (A, male Fakfak; B, female Arguni); C, R. ornata sp. n. (male South N.G.); D–E, R. variata variata ssp. n. male (Nabire); F–G, R. nigropunctata sp. n., male (Mt. Nok); H–I, R. camela sp. n., male (Sorong); J–K, R. arfak sp. n., (J, female Siwi; K, male Koffo); L, R. genyem sp. n. female (Genyem); M, R. variata brevis ssp. n. male (Freeport); N, Haudrhytidaspis fusca (Karny, 1911) comb. nov. female (type).—Habitus dorsal view (A, C, E–F, H, K, M–N) and lateral view (B, D, G, I–J, L). Scales 10 mm.

FIGURE 2. Face of A–B, Rhytidaspis variata meja ssp. n. (A, male Tafelberg; B, female Dorey); C–E, R. arfak sp. n. (C, male Koffo; D, male Siwi; E, female Siwi); F, R. variata brevis ssp. n. male (Freeport); G, R. variata variata ssp. n. (male Nabire); H–J, R. picta Redt., 1891 (H, male Fakfak; I, female Arguni; J, male Bundi); K–M, R. ornata sp. n. (K, male South N.G.; L, male Mindiptana; M, male Tiel); N, R. camela sp. n. (male Sorong); O, R. nigropunctata sp. n. (male Mt. Nok); P, R. genyem sp. n. (female Genyem); Q, Haudrhytidaspis fusca (Karny, 1911) comb. nov. (female type). Scales 10 mm. The arrows point at the sub-ocular carina, present in Rhytidaspis, absent in Haudrhytidaspis.

FIGURE 3. Left and right male tegmina of A–D, Rhytidaspis picta Redt., 1891 (A–B, Fakfak; C–D, Banda); E–F, R. arfak sp. n. (Koffo); G–H, R. variata meja ssp. n. (Tafelberg); I–J, R. variata variata ssp. n. (Nabire); K–L, R. variata brevis ssp. n. (Freeport); M–N, R. camela sp. n. (Sorong); O–P, R. nigropunctata sp. n. (Mt. Nok); Q–R, R. arfak sp. n. (Siwi); S–X, R. ornata sp. n. (S–T, South N.G.; U–V, Mindiptana; W–X, Tiel); Y, stridulatory file on underside of left tegmen of R. picta Redt., 1891 (Fakfak). Scales 5 mm, in Y 1 mm.

FIGURE 4. Stridulatory file on underside of left tegmen of A, Rhytidaspis arfak sp. n. (Koffo); B–C, R. camela sp. n. (B, Sorong; C, Komara); D, R. variata meja ssp. n. (Tafelberg); E, R. variata variata ssp. n. (Nabire); F, R. variata brevis ssp. n. (Freeport); G, R. nigropunctata sp. n. (Mt. Nok); H–J, R. ornata sp. n. (H, South N.G.; I, Mindiptana; J, Tiel).

FIGURE 5. Male abdominal apex with cerci in dorsal view: A, Rhytidaspis picta Redt., 1891 (Fakfak); B, R. arfak sp. n. (Koffo); C, R. nigropunctata sp. n. (Mt. Nok); D–E, R. camela sp. n. (D, Sorong; E, Komara); F–G, R. variata meja ssp. n. (F, Dorey; G, Tafelberg); H–I, R. ornata sp. n. (H, Tiel; I, Mindiptana); J, R. variata brevis ssp. n. (Freeport); K, R. variata variata ssp. n. (Nabire).

FIGURE 6. Male abdominal apex in apical view: A, Rhytidaspis picta Redt., 1891 (Fakfak); B, R. arfak sp. n. (Koffo); C–D, R. camela sp. n. (C, Komara; D, Sorong); E–F, R. ornata sp. n. (E, South N.G.; F, Mindiptana); G, R. variata brevis ssp. n. (Freeport); H, R. nigropunctata sp. n. (Mt. Nok); I, R. ornata sp. n. (Tiel).—Abbreviations: at apical lobe of tenth tergite, dc dorsal apical lobe of cercus, e epiproct, pa paraproct, pp projection of paraproct, vc ventral apical lobe of cercus.

FIGURE 7. Male abdominal apex or only cercus in lateral and ventral view: A–C, Rhytidaspis arfak sp. n. (A–B, Siwi; C, Koffo); D–E, R. picta Redt., 1891 (Fakfak); F–G, R. ornata sp. n. (South N.G.); H–J, R. camela sp. n. (H, Komara; I–J, Sorong); K–M, R. ornata sp. n. (K–L, South N.G.; M, Tiel); N, R. variata variata ssp. n. (Nabire); O–Q, R. variata meja ssp. n. (O–P, Tafelberg; Q, Dorey); R–S, R. variata brevis ssp. n. (Freeport); T–U, R. nigropunctata sp. n. (Mt. Nok).—Ventral view (A, E–I, K, P, S, U), oblique lateral view (B–D, J, L–O, Q–R, T).

FIGURE 8. Male titillators in lateral or oblique lateral view (A in internal, B in external view; E left in internal, right in top view): A–C, Rhytidaspis arfak sp. n. (A–B, Siwi; C, Koffo); D, R. picta Redt., 1891 (Fakfak); E–F, R. camela sp. n. (E, Sorong; F, Komara); G, R. nigropunctata sp. n. (Mt. Nok); H–J, R. ornata sp. n. (H, Tiel; I, Mindiptana; J, South N.G.); K, R. variata variata ssp. n. (Nabire); L–M, R. variata meja ssp. n. (L, Tafelberg; M, Dorey); N, R. variata brevis ssp. n. (Freeport).—Abbreviations: ap apical projection of granular sac, as oval apico-lateral sclerite, at apical branch of titillator, ea elongate apical sclerite, gs granular sac on top of U-shaped fold, ms basal muscular sac, ti base of titillator.—Tip of titillator is on top (A–F, H–K); tip is right (G, L–N).

FIGURE 9. Male titillators in ventro-apical view (tip on top): A, Rhytidaspis picta Redt., 1891 (Fakfak); B, R. arfak sp. n. (Koffo); C, R. variata meja ssp. n. (Tafelberg); D, R. variata brevis ssp. n. (Freeport); E, R. variata variata ssp. n. (Nabire); F, R. camela sp. n. (Sorong); G, R. nigropunctata sp. n. (Mt. Nok); H–I, R. ornata sp. n. (H, Mindiptana; I, South N.G.).—Abbreviation: fc flexible connection between both titillators.