Rhytidaspis variata brevis, Ingrisch, 2019

Ingrisch, Sigfrid, 2019, Revision of the genus Rhytidaspis Redtenbacher, 1891 including the description of a new genus Haudrhytidaspis gen. nov. (Orthoptera: Tettigoniidae: Conocephalinae), Zootaxa 4661 (2), pp. 343-370 : 354-358

Rhytidaspis variata brevis ssp. n.

Figs. 1M , 2F , 3 K–L, 4F, 5J, 6G, 7R–S, 8N, 9D

Holotype (male): Indonesia, Papua: Freeport [4°5’S, 137°7’E], 26–28.iii. (year not recorded, label reads /174), leg. Xavier Miçonne—(Brussels, ISNB).

Diagnosis. R. v. brevis ssp. n. differs from both other subspecies of R. variata by the dorsal apical lobe of the male cercus that has the proximal angle prolonged into a compressed triangular lobe and the distal end terminating into a narrowly rounded lobe while the proximal end is finger-shaped and the distal end about angularly projecting in both other subspecies. Moreover, the stridulatory file on the underside of the left tegmen is with 2.4 mm markedly shorter in R. v. brevis than in R. v. meja (2.6 mm) and R. v. variata (2.8 mm), while the density of teeth in the middle of the stridulatory file is nearly identical with that in R. v. variata but less dense than in R. v. meja .

Description of male. Tegmen behind stridulatory area with converging lateral margins, apex rounded. Stridulatory file on underside of left tegmen 2.4 mm long, with about 82 teeth, in middle with 27.5 teeth per 1 mm, toward end teeth become very small and dense ( Fig. 4F ). Mirror on right tegmen with nearly parallel fore and hind margins, internal margin strongly oblique, 1.61 mm wide, 1.05 mm long ( Fig. 3L ).

Cerci with dorsal apical lobe running oblique and parallel along inner margin of cercus trunk to nearly basal third of cercus; on apical side surpassing cercus trunk by a rounded lobe; margin of lobe very faintly wavy, the basal most area is separated by a v-shaped incision from the remainder, is dorsoventrally widened and terminates into a compressed-rounded lobe ( Fig. 5J ); the ventral apical lobe is dorsoventrally compressed with dorsal side convex and ventral side concave; on ventral side it is separated from the cercus trunk by a transverse bulge; apical margin down-curved, sub-truncate but little concave in middle; a strong curved acute tooth inserted at inner half of apical margin pointing ventrad ( Fig. 6G ).

Titillators with granular extension above the U-shaped curvature with margin in middle sub-straight to slightly concave, at proximal end rounded, and distally prolonged into a short, in lateral view triangular projection; separated by a long arm from little curved tip of titillator, which is only in apical area covered by spinules and a few larger teeth ( Figs. 8N , 9D ).

Coloration: Light to medium brown; pronotum and abdominal tergites of somewhat darker color ( Fig. 1M ). Antennal scrobae, scapus and pedicellus black, flagellum yellowish ( Fig. 2F ). Meso- and metapleura and lateral areas of abdominal tergites black. Apical cones of tenth abdominal tergites and cerci yellow. Tegmen bright yellow with black dots, stridulatory area semi-transparent. Legs yellow; femora with black transverse strokes; hind femur in basal area medium to dark red brown. Thoracic sternites and parts of coxae of all legs dark brown or black. Male subgenital plate black.

Measurements (1 male).—Body: 32; pronotum: 11; tegmen: 8; hind femur: 22.5; anterior femur: 11 mm.

Etymology. The name of the new subspecies refers to the short stridulatory file compared to both other subspecies; from Latin brevis short (Maskulinum and Femininum).

Figures

FIGURE 1. A–B, Rhytidaspis picta Redt., 1891 (A, male Fakfak; B, female Arguni); C, R. ornata sp. n. (male South N.G.); D–E, R. variata variata ssp. n. male (Nabire); F–G, R. nigropunctata sp. n., male (Mt. Nok); H–I, R. camela sp. n., male (Sorong); J–K, R. arfak sp. n., (J, female Siwi; K, male Koffo); L, R. genyem sp. n. female (Genyem); M, R. variata brevis ssp. n. male (Freeport); N, Haudrhytidaspis fusca (Karny, 1911) comb. nov. female (type).—Habitus dorsal view (A, C, E–F, H, K, M–N) and lateral view (B, D, G, I–J, L). Scales 10 mm.

FIGURE 2. Face of A–B, Rhytidaspis variata meja ssp. n. (A, male Tafelberg; B, female Dorey); C–E, R. arfak sp. n. (C, male Koffo; D, male Siwi; E, female Siwi); F, R. variata brevis ssp. n. male (Freeport); G, R. variata variata ssp. n. (male Nabire); H–J, R. picta Redt., 1891 (H, male Fakfak; I, female Arguni; J, male Bundi); K–M, R. ornata sp. n. (K, male South N.G.; L, male Mindiptana; M, male Tiel); N, R. camela sp. n. (male Sorong); O, R. nigropunctata sp. n. (male Mt. Nok); P, R. genyem sp. n. (female Genyem); Q, Haudrhytidaspis fusca (Karny, 1911) comb. nov. (female type). Scales 10 mm. The arrows point at the sub-ocular carina, present in Rhytidaspis, absent in Haudrhytidaspis.

FIGURE 3. Left and right male tegmina of A–D, Rhytidaspis picta Redt., 1891 (A–B, Fakfak; C–D, Banda); E–F, R. arfak sp. n. (Koffo); G–H, R. variata meja ssp. n. (Tafelberg); I–J, R. variata variata ssp. n. (Nabire); K–L, R. variata brevis ssp. n. (Freeport); M–N, R. camela sp. n. (Sorong); O–P, R. nigropunctata sp. n. (Mt. Nok); Q–R, R. arfak sp. n. (Siwi); S–X, R. ornata sp. n. (S–T, South N.G.; U–V, Mindiptana; W–X, Tiel); Y, stridulatory file on underside of left tegmen of R. picta Redt., 1891 (Fakfak). Scales 5 mm, in Y 1 mm.

FIGURE 4. Stridulatory file on underside of left tegmen of A, Rhytidaspis arfak sp. n. (Koffo); B–C, R. camela sp. n. (B, Sorong; C, Komara); D, R. variata meja ssp. n. (Tafelberg); E, R. variata variata ssp. n. (Nabire); F, R. variata brevis ssp. n. (Freeport); G, R. nigropunctata sp. n. (Mt. Nok); H–J, R. ornata sp. n. (H, South N.G.; I, Mindiptana; J, Tiel).

FIGURE 5. Male abdominal apex with cerci in dorsal view: A, Rhytidaspis picta Redt., 1891 (Fakfak); B, R. arfak sp. n. (Koffo); C, R. nigropunctata sp. n. (Mt. Nok); D–E, R. camela sp. n. (D, Sorong; E, Komara); F–G, R. variata meja ssp. n. (F, Dorey; G, Tafelberg); H–I, R. ornata sp. n. (H, Tiel; I, Mindiptana); J, R. variata brevis ssp. n. (Freeport); K, R. variata variata ssp. n. (Nabire).

FIGURE 6. Male abdominal apex in apical view: A, Rhytidaspis picta Redt., 1891 (Fakfak); B, R. arfak sp. n. (Koffo); C–D, R. camela sp. n. (C, Komara; D, Sorong); E–F, R. ornata sp. n. (E, South N.G.; F, Mindiptana); G, R. variata brevis ssp. n. (Freeport); H, R. nigropunctata sp. n. (Mt. Nok); I, R. ornata sp. n. (Tiel).—Abbreviations: at apical lobe of tenth tergite, dc dorsal apical lobe of cercus, e epiproct, pa paraproct, pp projection of paraproct, vc ventral apical lobe of cercus.

FIGURE 8. Male titillators in lateral or oblique lateral view (A in internal, B in external view; E left in internal, right in top view): A–C, Rhytidaspis arfak sp. n. (A–B, Siwi; C, Koffo); D, R. picta Redt., 1891 (Fakfak); E–F, R. camela sp. n. (E, Sorong; F, Komara); G, R. nigropunctata sp. n. (Mt. Nok); H–J, R. ornata sp. n. (H, Tiel; I, Mindiptana; J, South N.G.); K, R. variata variata ssp. n. (Nabire); L–M, R. variata meja ssp. n. (L, Tafelberg; M, Dorey); N, R. variata brevis ssp. n. (Freeport).—Abbreviations: ap apical projection of granular sac, as oval apico-lateral sclerite, at apical branch of titillator, ea elongate apical sclerite, gs granular sac on top of U-shaped fold, ms basal muscular sac, ti base of titillator.—Tip of titillator is on top (A–F, H–K); tip is right (G, L–N).

FIGURE 9. Male titillators in ventro-apical view (tip on top): A, Rhytidaspis picta Redt., 1891 (Fakfak); B, R. arfak sp. n. (Koffo); C, R. variata meja ssp. n. (Tafelberg); D, R. variata brevis ssp. n. (Freeport); E, R. variata variata ssp. n. (Nabire); F, R. camela sp. n. (Sorong); G, R. nigropunctata sp. n. (Mt. Nok); H–I, R. ornata sp. n. (H, Mindiptana; I, South N.G.).—Abbreviation: fc flexible connection between both titillators.