Diplometopon Nikolski, 1907: 276

GANS, CARL, 2005, Checklist And Bibliography Of The Amphisbaenia Of The World, Bulletin of the American Museum of Natural History 2005 (289), pp. 1-130 : 42-43

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scientific name	Diplometopon Nikolski, 1907: 276
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Diplometopon Nikolski, 1907: 276 . Type species: Diplometopon zarudnyi , by monotypy.

See discussion in Gans (1960: 178).

Diplometopon zarudnyi Nikolski

Diplometopon zarudnyi Nikolski, 1907: 277 . TYPE LOCALITY: Nasrie in Arabistano ,... in Persia australooccidentali’’ = Nasrie in southwestern Iran on the Karun River. Holotype: ZIL 10341.

Pachycamalus zaroudnyi G. Boulenger, 1920: 352. Emendation: Collector’s name spelled ‘‘Zaroudny’’ in title of Nikolski’s paper.

DISCUSSION OF FORM: Gans (1960: 178). See also AlBadry and AlSidirawi (1976, blood), AlGhandoor (1992, reproduction), AlJohany (1999, metabolism), AlNassar (1976, osteology, gut histology), AlSadoon (1986, metabolism; 1988, distribution), Al Sadoon et al. (1991, ecology), AlThani and ElSherif (1996, digestive tract histology), AlYousif and AlShawa (1998, parasites), Anderson (1963, 1967, 1968, 1974, 1979, distribution), Arnold (1984a, hemipenes), Branch (1980, chromosomes), Cloudsley Thompson (1979, physiology), ElAssy and AlNassar (1976, anatomy), Gallagher (1971, distribution), Gans (1971c, list), Gans and Wever (1975, anatomy), Gasperetti (1967, distribution), Haas and Werner (1969, distribution), Hellyer (1994), Henson (1974, anatomy), Khalaf (1960, distribution), Lanza (1983, distribution), Leviton and Anderson (1967, distribution), Mahdi and Georg (1969, distribution), Mertens (1959, 1960, general), Miller (1966a, anatomy), Niazi (1979, ecology), Tuck (1971, list), and Yousif and Al Shawa (1998, parasites).

RANGE: Extreme western Iran, southern Iraq, Kuwait, the northern half of Saudi Arabia, Trucial Oman.

Genus Pachycalamus Günther

Pachycalamus C.A.L.G. Günther, 1881: 462 . Type species: Pachycalamus brevis , by monotypy.

See discussion in Gans (1960: 178).

Pachycalamus brevis Günther Pachycalamus brevis C.A.L.G. Günther, 1881: 462 . TYPE LOCALITY: Socotra Island. Syntypes: BMNH 81.3.5.6.:RR 1946.8.31 –81.3.5.11: RR1946.8–31.73, 1934.12.10.1:RR1946.8.31.78– 1934.12.10.4:RR1946.8.31.81 (possibly NMW 16681 as well).

DISCUSSION OF FORM: Loveridge (1941: 364), Gans (1960: 178). See also Boulenger (1903, 1920, distribution), Gans (1971c, list), Joger (2000), Kennedy (1937, distribution), and Peters (1882a).

RANGE: Socotra Island, Indian Ocean.

Genus Trogonophis Kaup

Trogonophis Kaup, 1830: 880 . Type species: Trogonophis wiegmanni , by monotypy.

Trogonophis Gray, 1844: 68 . Lapsus , hence not available (International Code of Zoological Nomenclature, 1961, art. 33b).

See discussion in Gans (1960: 178).

Trogonophis elegans ( Gervais) Amphisbaena elegans Gervais, 1835: 113 . TYPE LOCALITY: ‘‘ Alger et à Tanger’ ’; restricted to Tangier (Gans and Pasteur, 1962: 10). Lectotype: MHNP A3083 from Tangier. Lectoparatypes: Apparently lost (cf. Gans and Pasteur, 1962).

Trogonophis wiegmanni maroccana Werner, 1931b: 280 . TYPE LOCALITY: ‘‘Chella, Rabat, Marokko’’. Holotype: MCZ 29668. Paratypes: MCZ 29669 (Djebel Taghat, Fez). MCZ 29670 ( Agadir, Morocco).

DISCUSSION OF FORMS: See under the nominate race.

RANGE: Western Morocco to the Souss.

Trogonophis wiegmanni Kaup

Trogonophis wiegmanni Kaup, 1830: 880 . TYPE LOCALITY: Unknown; restricted to ‘‘Algérie’’

( Pasteur and Bons, 1960: 79). Syntypes: ‘‘Zool.

Inst. Giessen’’ (Kaup, 1831); destroyed (W.J.

Schmidt, in litt.).

DISCUSSION OF FORMS: Loveridge (1941: 360), Gans (1960: 178). Pasteur and Bons (1960: 79) commented on the status of races. See also Anderson (1892, distribution), Angel and RochonDuvigneaud (1942b, anatomy), Baker (1987, parasites), Barbier (1906, list), Bellairs (1950, anatomy), Bellairs and Boyd (1947, anatomy), Blackburn (1985, biology), Boettger (1881, 1883, 1885a, distribution), Böhme (1970, general), Bons and Girot (1962, identification), Bons and Saint Girons (1963, biology), Boulenger (1891c, 1905a, distribution), Burlet (1934, anatomy), Busack (1976, list), Capula (1989, identification), Chabanaud (1916, distribution), Doumergue (1900, distribution), Dufaure and Saint Girons (1984, anatomy), Fischer (1889), Flower (1925, longevity), Fürbringer (1870, 1900, anatomy), Gabe and Saint Girons (1971, 1972, 1976 anatomy), Gabe et al (1964), Gans and Pasteur (1962, distribution), Gatten and McClung (1981, behavior), Gervais (1848b, distribution; 1853, anatomy), Guibé (1954, list), Hediger (1937a, 1937b, distribution), Jacobshagen (1920, anatomy), Kaup (1830), Lakjer (1927, anatomy), Le Cerf (1909), Makino (1951, chromosomes), Matthey (1931, 1932a, 1932c, 1951b, chromosomes), Mertens (1915), Olivier (1894, 1896, distribution), Pellegrin (1912a, distribution), Pratt (1948, anatomy), Rathke (1853), Remane (1936, anatomy), Richter (1933, anatomy), Rieppel (1979, 1988a, 1996, 1971, anatomy), RochonDuvigneaud (1954, anatomy), Saint Girons (1953, 1961, 1972, 1989, anatomy), Saint Girons and Joly (1975, anatomy), Schleich et al. (1996, identification), Smalian (1884a, anatomy), Stemmler (1970, distribution), Verrier (1932, 1933, anatomy), Versluys (1912, anatomy), Werner (1929a, 1931b), Woerdeman (1921b), Zangerl (1944, 1945, anatomy), and Zulueta (1909, distribution).

RANGE: Eastern Morocco, northern Algeria, western Tunisia (specimens in Museo Civico di Storia Naturale of Genoa and the Zoologische Staatssammlung, Munich).

References

Loveridge, A. 1941. Revision of the African lizards of the family Amphisbaenidae. Bulletin of the Museum of Comparative Zoology 87 (5): 353 - 451. [all]

Mahdi, N., and P. V. Georg. 1969. A systematic list of the vertebrates of Iraq. Iraq Natural History Museum Publication 26: 1 - 104. [p. 30; Amphisbaenidae, Amphisbaeninae, Diplometopon zarudnyi]

Makino, S. 1951. An atlas of the chromosome numbers in animals, 2 nd ed. Ames: Iowa State College Press. [p. 244; Amphisbaenidae, Rhineura floridana 46 S, Trogonophis wiegmanni 36 S]

Matthey, R. 1931. Les chromosomes des amphisbeniens acrodontes (Trogonophis wiegmanni, Kaup). Bulletin de la Societe Vaudaume de Sciences Naturelles 57 (227): 313. [all]

Matthey, R. 1932 a. Les chromosomes de l'amphisbenien acrodonte Trogonophis wiegmanni Kaup. Archives de Zoologie Experimentale et Generale, volume jubilaire, 74: 193 - 203. [p. 196]

Matthey, R. 1932 c. Nouvelle contribution a l'etude des chromosomes chez les Sauriens. Bulletin de la Societe Vaudaume de Sciences Naturelles 57 (231): 587 - 588. [p. 587, Rhineura floridana, Trogonophis wiegmanni; p. 588, Amphisbaenidae]

Matthey, R. 1951 b. Systematique et criteres cytologique. Les chromosomes des Blanus cinereus Vand. (Lacertilia - Amphisbaenidae) et la meiose de Leucophaea maderae (Blattaria - Blaberidae). Bulletin de la Societe Vaudaume de Sciences Naturelles 65 (279): 111 - 120. [all; Blanus cinereus from Rabat (= Blanus tingitanus?), Rhineura floridana, Trogonophis wiegmanni]

Mertens, R. 1915. Eine Amphisbaenide im Terrarium. Wochenschrift der Aquarien und Terrarien Kunde 1915: 65 - 66. [all; Trogonophis wiegmanni]

Mertens, R. 1959. La vie des amphibiens et reptiles. Paris: Horizons de France. [p. 28, 149; Blanus, Diplometopon]

Miller, M. R. 1966 a. The cochlear duct of lizards. Proceedings of the California Academy of Sciences, ser. 4, 23: 255 - 359. [p. 294; Amphisbaena fuliginosa, Bipes biporus, Diplometopon zarundnyi, Rhineura floridana]

Niazi, A. D. 1979. The Arabian worm lizard: its ecology, morphology and phylogeny with description of a new species. Iraq Journal of Biological Sciences 7 (1): 3 - 29. [all; Diplometopon shuebaibi, D. zarudniyi]

Nikolski, A. M. 1907. Reptiles et amphibiens, recueillis par Mr. N. A. Zaroudny en Perse en 1903 - 1904. Annales de le Museum de Zoologie, Academie Imperial de Science de St. Petersbourg 10 (3, 4): 260 - 301. [pp. 276 - 280]

Olivier, E. 1894. Herpetologie Algerienne ou catalogue raissonne des reptiles et des batraciens observes jusqu'a ce jour en Algerie. Memoirs de la Societe Zoologique de France 7 (1): 98 - 131. [pp. 105, 117; Blanus cinereus, Trogonophis wiegmanni]

Olivier, E. 1896. Materiaux pour la faune de la Tunisie. I. Catalogue de reptiles. Revues de Sciences Bourbonnais Centre France (Moulins) 9: 117 - 128. [pp. 122 - 123; Trogonophis wiegmanni]

Pasteur, G., and J. Bons. 1960. Catalogue des reptiles actuels du Maroc. Revision de formes d'Afrique, d'Europe et d'Asie. Travaux Institute de Science de Cherifien, Ser. Zoologie, 21: 1 - 132. [pp. 21, 77 - 79]

Pellegrin, J. 1912 a. Reptiles, batraciens, et poissons du Maroc (Mission de Mme. Camille du Gast). Bulletin de la Societe Zoologique de France 37 (7): 255 - 262. [p. 256; Trogonophis wiegmanni]

Peters, W. C. H. 1882 a. Die von Herrn Dr. E. Riebeck auf Socotra gesammelten Reptilien. Sitztungsberichte der Gesellschaft Naturforschen- der Freunde zu Berlin 1882: 42 - 46. [p. 46; Pachycalamus brevis]

Pratt, C. W. M. 1948. The morphology of the ethmoidal region of Sphenodon and lizards. Proceedings of the Zoological Society of London 118: 171 - 201. [pp. 172, 182, 196, Amphisbaena alba, Monopeltis, Trogonophis wiegmanni]

Rathke, H. 1853. Ueber den Bau und die Entwicklung des Brustbeins der Saurier. Konigsberg: Albertus-Universitat. [pp. 1, 3; Amphisbaena alba, Trogonophis]

Remane, A. 1936. Skelettsystem. I. Wirbelsaule und ihre Abkommlinge. In L. Bolk, E. Goppert, E. Kallius, and W. Lubosch (editors), Handbuch der vergleichenden Anatomie der Wirbeltiere 4: 1 - 206. Berlin: Urban und Schwarzenberg. [p. 112, Amphisbaena; pp. 158, 178, Chirotes canaliculatus; p. 180, Blanus cinereus, Trogonophis wiegmanni; p. 181, Blanus cinereus, B. strauchi, Trogonophis wiegmanni]

Richter, H. 1933. Das Zungenbein und seine Muskulatur bei den Lacertilia vera. Jenaische Zeitschrift der Naturwissenschaften, n. ser., 66 (59): 395 - 480. [Amphisbaena darwini, A. fuliginosa, Monopeltis semipunctata, Trogonophis wiegmanni]

Rieppel, O. 1971. Zum Verhalten von Trogonophis wiegmanni elegans (Gervais, 1835). Aquaterra 8 (4): 44 - 48. [all]

Rieppel, O. 1979. The external jaw adductor of amphisbaenids (Reptilia: Amphisbaenia). Revue Suisse de Zoologie 86 (4): 867 - 876. [used Amphisbaena alba, A. fuliginosa, Trogonophis wiegmanni elegans, T. w. wiegmanni. Situation is quite different from snakes]

Rieppel, O. 1988 a. The classification of the Squamata. In M. J. Benton (editor), The phylogeny and classification of the Tetrapods, vol. 1: Amphibians, reptiles, birds. Systematics Society Special Volume 35 A: 261 - 293. Oxford: Clarendon Press. [pp. 261 - 267, 272, 277, 278, 280 - 282; Trogonophis]

Rieppel, O. 1996. Miniaturization in tetrapods: consequences for skull morphology. Symposia of the Zoological Society of London 69: 47 - 61. [pp. 50 - 51, 53, 57; fig. 1; Amphisbaena alba mislabeled as Trogonophis wiegmanni]

Rochon-Duvigneaud, A. 1954. L'oeil des vertebres. In P. - P. Grasse (editor), Traite de Zoologie 12: 333 - 452. Anatomie, systematique, biologie. Paris: Masson et Cie. [p. 437; Trogonophis]

Saint Girons, H. 1953. Influence de la mise en eau du barrage de Bin el Ouidane sur les vertebres terrestres. Bulletin de la Societe Scientifique et Naturaliste de Maroc 33 (2): 75 - 80. [Blanus cinereus, Trogonophis]

Saint Girons, H. 1961. Particularites anatomiques et histologiques de l'hypophyse chez la Squamata. Archives de Biologie 72 (2): 211 - 299. [pp. 247 - 250; fig. 16; Blanus cinereus, Trogonophis wiegmanni]

Saint Girons, H. 1971. Quelques facteurs du rythme circadien d'activite chez les lepidosauriens (reptiles). Bulletin de la Societe Zoologique de France 96: 317 - 330. [p. 330]

Saint Girons, H. 1972. Morphologie comparee du segment sexuel du rein des squamates (Reptilia). Archives d'Anatomie Microscopique et Morphologie Experimentale 61 (3): 243 - 266. [pp. 248, 251, 254, 260; Amphisbaena alba, Trogonophis wiegmanni]

Saint Girons, H., and J. Joly. 1975. Histologie et ultrastructure de la glande nasale du lacertilien Lacerta muralis et de l'amphisbenien Trogonophis wiegmanni (Reptilia, Lacertidae et Trogonophidae). Archives de Biologie (Bruxelles) 86: 97 - 126. [all]

Saint Girons, H. 1989. Les glandes cephaliques exocrines des reptiles. II. Considerations fonctionelles et evolutives. Donnees anatomiques et histologiques. Annales de Sciences Naturelles, Zoologie, Paris, ser. 13, 10: 1 - 17. [pp. 5 - 6; Trogonophis]

Schleich, H. H., W. Kastle, and K. Kabisch. 1996. Amphibians and reptiles of North Africa: biology, systematics, field guide. Koenigstein: Koeltz Scientific Books. [Blanus cinereus, Trogonophis]

Smalian, C. 1884 a. Beitrage zur Anatomie der Amphisbaeniden. Zeitschrift der Wissenschaften, Zoologie 42: 126 - 202. [all; pls. 5, 6; Amphisbaena fuliginosa, Anops kingii, Blanus cinereus, Trogonophis wiegmanni]

Stemmler, O. 1970. Beobachtungen an marokkanischen Schachbrettschleichen, Trogonophis wiegmanni Kaup 1830 (Amphisbaenia, Trogonophidae). Aquarien und Terrarien 17 (10): 343 - 347. [all]

Tuck, R. G., Jr. 1971. Amphibians and reptiles from Iran in the United States National Museum Collection. Bulletin of the Maryland Herpetological Society 7: 48 - 86. [p. 60; map, p. 72; Diplometopon zarudnyi,]