Stamnodes coenonymphata ( Hulst, 1900 )

Matson, Tanner A., 2023, A review of Mexican Stamnodes (Lepidoptera: Geometridae) with the description of 16 new species, European Journal of Taxonomy 911, pp. 1-79 : 51-52

Stamnodes coenonymphata ( Hulst, 1900)

Figs 25 , 91 , 94–95

Coenocalpe coenonymphata Hulst, 1900: 104 . Type locality: Pasadena, California; Yosemite Valley, California, USA. [AMNH].

Stamnodes coenonymhata prunata Wright, 1924: 92–93 . Type locality: San Diego, California, USA. [SDNHM, MCZ].

Stamnodes coenonymphata brunneata Wright, 1924: 93 . Type locality: San Diego, California, USA. [SDNHM, MCZ].

Stamnodes coenonymphata pallidata Wright, 1924: 93 . Type locality: San Diego, California, USA. [SDNHM, MCZ].

Stamnodes coenonymphata wrightii Cassino & Swett, 1925: 38 . Type locality: San Diego, California, USA. [Not found in MCZ].

Coenocalpe coenonymphata – Dyar 1902: 285 (cat.).

Stamnodes coenonymphata – Pearsall 1909b: 366. — McDunnough 1938: 151 (checklist). — McFarland 1965: 62. — Ferguson 1983: 103 (checklist). — Furniss et al. 1988: 7–9. — Poole & Gentili 1996: 686 (checklist). — Brown & Bash 2000: 73. — Scoble 1999: 901 (catalogue). — Miller & Hammond 2003: 158. — Scoble & Hausmann 2007 (online catalogue). — Pohl et al. 2016: 448 (checklist). — Rajaei et al. 2022 (online catalogue). — Albu & Albu 2023: 55 (survey).

Diagnostic remarks

Stamnodes coenonymphata is highly variable in colour, especially the hindwing underside.The transverse medial band of the hindwing underside common to many Cercocarpus -feeding Stamnodes is generally white in S. coenonymphata with the scales basad and distad greatly differing in colour. The transverse band is more basally positioned than in congeners and has a more acute angle as it extends basad toward the margins. This species is most often confused with S. costimacula , S. annellata , and S. cassinoi .

Distribution

Mexico: Stamnodes coenonymphata is found in the chaparral associations, foothills, canyons, and Pacific coastal scrub communities of northwestern Baja California. USA: the core of this species’ range lies in California.

Biology

The peak flight of S. coenonymphata is January to March with larvae to follow in the spring and early summer. Larvae ( Fig. 91 ) are hosted by birch leaf mountain mahogany ( Cercocarpus betuloides Nutt. ) ( McFarland 1965).

Molecular characterization

This species is represented in BOLD as BIN: BOLD:AAF2593 (n = 26). At present, the average pairwise intraspecific distance is 0.82%, the pairwise maximum intraspecific distance is 1.93%, and the distance to the nearest adjacent interspecific neighbour, Stamnodes formosata (n = 20), is about 3%.

Figures

Figs 19–30. Adult Stamnodes Guenée, [1858], dorsal (a) and ventral (b) views. 19. S. saltillo sp. nov., holotype, ♂ (CNIN). 20. S. formosata (Strecker, 1878) (USNMENT01771263). 21. S. lampra Rindge, 1878, paratype (USNMENT01771264). 22. S. franckata (Pearsall, 1909) (USNMENT01771255). 23. Stamnodes sp. (?undescribed; Mexico: Nuevo León) (USNMENT01523823). 24. S. patamon (Druce, 1893) (CNIN). 25. S. coenonymphata (Hulst, 1900) (TAM). 26. S. catarina sp. nov., holotype, ♂ (AMNH_IZC 00352926). 27. S. churro sp. nov., holotype, ♀ (AMNH_IZC 00352920). 28. S. ferropulvisa sp. nov., holotype, ♀ (USNMENT01523824). 29. S. calcarea sp. nov., holotype, ♂ (CMNH). 30. S. ululata Pearsall, 1912 (USNMENT01771256). Scale bar = 1 cm.

Figs 90–93. Final instar larval variation of North American species of Stamnodes Guenée, [1858]. 90. S. albiapicata Grossbeck, 1910, collected from Pholistoma membranaceum (Benth.) Constance; USA: California: San Diego Co. (2017C29); photo credit Moria L. Robinson. 91. S. coenonymphata (Hulst, 1900) collected from Cercocarpus betuloides Nutt.; USA: California: Sonoma Co. (2018E171). 92. S. formosata (Strecker, 1878) collected from Cercocarpus montanus Raf.; USA: Arizona: Cochise Co. (2014K98), photo credit David L. Wagner. 93. S. fergusoni Matson & Wagner, 2020, collected from Salvia pinguifolia (Fernald) Wooton & Standl.; USA: New Mexico: Lincoln Co. (2018J32), photo credit David L. Wagner (Matson & Wagner 2020).

Fig. 94. COI neighbour-joining tree for Mexican Stamnodes Guenée, [1858] using default Kimura-2P model in the Barcode of Life Project (BOLD). Taxon names followed by BOLD Process ID.

Fig. 95. Biogeographic associations of Mexican Stamnodes Guenée, [1858]. (*) denotes species found in two or more provinces. Underlying biogeographic provincial map taken from Morrone (2020: fig. 4.5) with permission to reproduce this figure granted by author Juan J. Morrone.