Euthria walaszczyki, Harzhauser & Landau, 2024

Harzhauser, Mathias & Landau, Bernard M., 2024, The Colubrariidae, Eosiphonidae, Melongenidae, Pisaniidae, Prodotiidae and Tudiclidae (Gastropoda, Buccinoidea) of the Miocene Paratethys Sea, Zootaxa 5427 (1), pp. 1-110 : 67-69

Euthria walaszczyki nov. sp.

Figs 34B , 35A–C

[ Fusus ] corneus — Naumann 1852: plate captions, pl. 70, fig. 8 [non Euthria cornea ( Linnaeus, 1758) ].

Fusus corneus Linn. —Hoernes 1853: 280, pl. 31, figs 3a–b [non Euthria cornea ( Linnaeus, 1758) ].

Fusus corneus Linné — Neugeboren 1854: 184 [non Euthria cornea ( Linnaeus, 1758) ].

[ Fusus ] corneus Brocc. — Naumann 1854: 1066 [non Euthria cornea ( Linnaeus, 1758) ].

Fusus (Euthria) corneus Linn. —Hoernes & Auinger 1890: 257 [non Euthria cornea ( Linnaeus, 1758) ].

E [uthria]. (E [uthria].) cornea (L.)— Sieber 1958: 150 [non Euthria cornea ( Linnaeus, 1758) ].

Type material. Holotype: NHMW 1851 /0002/0044, SL: 43.1 mm, MD: 19.8 mm, Grund ( Austria), illustrated in Hoernes (1853: pl. 31, fig. 3); Figs 35A 1 –A 2 . Paratypes: NHMW 1863 /0015/1270, SL: 39.6 mm , MD: 17.8 mm, Grund ( Austria), Figs 35B 1 –B 2 . NHMW 1869 /0001/0281, SL: 33.3 mm , MD: 14.5 mm, Grund ( Austria), Figs 35C 1 –C 2 . NHMW 1868 /0001/0223, SL: 16.7 mm , MD: 7.7 mm, Grund ( Austria), Figs 34B .

Additional material. 5 spec., NHMW 1868 /0001/0223, Grund ( Austria) .

Type locality. Grund ( Austria) .

Type stratum. Silt and clay of the Grund Formation.

Age. Middle Miocene, early Badenian (Langhian).

Etymology. In honor of Ireneusz Piotr Walaszczyk (Faculty of Geology, University of Warsaw, Poland) for his help searching for type material.

Diagnosis. Medium-sized, moderately slender fusiform shell with moderately high spire of convex whorls, early teleoconch whorls weakly shouldered just above suture, sculpture changing early in ontogeny from prominent axial and spiral sculpture to relatively smooth on third teleoconch whorl, last whorl evenly rounded, lacking subsutural ramp, base strongly constricted with moderately long, narrow siphonal canal.

Description. Medium-sized, moderately slender fusiform shell of up to seven teleoconch whorls; apical angle ~45°. Protoconch damaged in all available specimens; last protoconch whorl high, convex. Early teleoconch whorls slightly shouldered; periphery just above abapical suture. Relatively narrow axial ribs, separated by broad interspaces, weakening adapically, overrun by four primary spiral cords and delicate secondaries intercalated; ribs fading on third teleoconch whorl. Later teleoconch whorls convex, separated by moderately incised suture, bearing faint spiral threads, periphery below mid-whorl. Last whorl attaining ~65% of total height, with regularly convex periphery, base strongly constricted, fasciole indistinct; surface smooth except for weak spirals over fasciole.Aperture moderately wide, pyriform. Columella broadly excavated, angled at transition to siphonal canal often accentuated by small denticle. Columellar callus indistinct, weakly delimited from base, smooth; Anal canal moderately incised, accentuated by weak parietal swelling or denticle; no anal denticle. Outer lip thin at peristome, bearing 12–14 elongated denticles a short distance behind peristome that extend as fine lirae deep within aperture. Siphonal canal moderately long, narrow, deflected to the left, shallowly notched.

Discussion. This species was confused by Naumann (1852), Hoernes (1853) and Hoernes & Auinger (1890) with the Pleistocene and Recent Mediterranean Euthria cornea ( Linnaeus, 1758) . The extant species differs in its shorter spire and shouldered whorls with markedly concave subsutural ramp (see Brunetti & Della Bella 2016: figs 4A–D; Fraussen & Stahlschmidt 2017: pl. 4, figs 67–84). The Middle Miocene Paratethyan Euthria brunettii nov. sp. differs in its prominent axial ribs on the spire whorls that persist at least onto the penultimate whorl. Euthria perpiniana ( Fontannes, 1879) , from the Pliocene of the Mediterranean Sea, is very similar in general appearance and in the early disappearance of its axial sculpture but is larger (SL up to 55 mm), has less convex whorls with a distinct subsutural concavity, although weaker than in E. cornea , and more prominent spiral cords on early teleoconch whorls ( Fraussen & Stahlschmidt 2017: pl. 2, figs 40–48). The Pliocene Mediterranean Euthria plioelongata ( Sacco, 1904) is distinguished by its comparatively flat-sided spire whorls and axials that persist slightly longer, fading on the fourth teleoconch whorl (see Brunetti & Della Bella 2016: figs 2A–E; Fraussen & Stahlschmidt 2017: pl. 1, figs 26–33).

Paleoenvironment. At the locality Grund fossiliferous channel fills, which formed in middle to outer neritic environments bear allochthonous assemblages uniting coastal-mudflat faunas with inner neritic ones ( Zuschin et al. 2005; Roetzel 2009).

Distribution in Central Paratethys. Badenian (Middle Miocene): North Alpine Foreland Basin: Grund ( Austria) (Hoernes & Auinger 1890).

Figures

FIGURE 1. Measurements for shell morphology; AA: apical angle, SL: shell length, MD: maximum diameter, LWH: last whorl height, AH: aperture height, AL: aperture length, AW: aperture width.

FIGURE 2. Colubraria subobscura (Hoernes & Auinger, 1884). A1–A3. NHMW 1854/0035/0170, paralectotype, Lăpugiu de Sus (Romania). B1–B2. NHMW 1859/0037/0018a, paralectotype, Lăpugiu de Sus (Romania). C1–C3. NHMW 1865/0001/0183, paralectotype, Lăpugiu de Sus (Romania). D1–D2. NHMW 1859/0037/0018b, paralectotype, Lăpugiu de Sus (Romania). E1–E3. NHMW 1874/0025/0021, lectotype, Lăpugiu de Sus (Romania). F1–F2. NHMW 2023/0338/0001, paralectotype, Lăpugiu de Sus (Romania).

FIGURE 34.A. Euthria brunettii nov.sp., NHMW2023/0338/0007,paratype, Lăpugiu de Sus(Romania).B.Euthria walaszczyki nov. sp., NHMW 1868/0001/0223, Grund (Austria). C. Euthria depressospira Bandat, 1943, NHMW 2023/0338/0008, Lăpugiu de Sus (Romania). D. Euthria diluvii (Eichwald, 1830) NHMW 1860/0001/0224b, Boršov (= Porstendorf) (Czech Republic). E. Euthria fuscocingulata (Hoernes & Auinger, 1890). NHMW 2023/0338/0011, paralectotype, Lăpugiu de Sus (Romania). F. Euthria frausseni nov. sp., NHMW 2023/0338/0012, paratype, Lăpugiu de Sus (Romania). G. Euthria odiosa nov. sp. NHMW 2023/0348/0001, paratype, Steinebrunn (Austria). H. Euthria puschii (Andrzejowski, 1830), NHMW 2023/0350/0002, Bad Vöslau (Austria). I. Euthria stuetzii (Naumann, 1852), NHMW 1846/0037/0254, Enzesfeld or Gainfarn (Austria). J. Euthria subnodosa (Hoernes & Auinger, 1890), NHMW 1846/0037/0274, Steinebrunn and Gainfarn (Austria). K. Euthria yesimae nov. sp., RGM 794 572, paratype, Pınarlar Yaylası, Akpınar (Turkey).

FIGURE 35. Euthria walaszczyki nov. sp. A1–A2. NHMW 1851/0002/0044, holotype, Grund (Austria). B1–B2. NHMW 1863/0015/1270, paratype, Grund (Austria). C1–C2. NHMW 1869/0001/0281, paratype, Grund (Austria).