Aplus anatolicus, Harzhauser & Landau, 2024

Harzhauser, Mathias & Landau, Bernard M., 2024, The Colubrariidae, Eosiphonidae, Melongenidae, Pisaniidae, Prodotiidae and Tudiclidae (Gastropoda, Buccinoidea) of the Miocene Paratethys Sea, Zootaxa 5427 (1), pp. 1-110 : 40-42

Aplus anatolicus nov. sp.

Figs 15E , 21A–B

Anna exsculpta (Dujardin) — Landau et al. 2013: 168, pl. 25, figs 10–11, pl. 64, fig. 8 [non Aplus exsculptus ( Dujardin, 1837) ].

Type material. Holotype: NHMW 1847 /0058/0889, SL: 18.6 mm, MD: 10.1 mm, Seyithasan ( Turkey), Serravallian (Middle Miocene), illustrated in Landau et al. (2013: pl. 21, fig. 10), Figs 21A 1 –A 2 . Paratypes: NHMW 1847 /0058/0979, SL: 18.6 mm , MD: 8.7 mm, Seyithasan ( Turkey), Serravallian (Middle Miocene), illustrated in Landau et al. (2013: pl. 21, fig. 11), Figs 21B 1 –B 3 . Paratypes. Additional paratypes: NHMW 1847 /0058/1694/1,

RGM 783 889/1, Akboğaz at Lale River; NHMW 1847/0058/0889/1, 1847/0058/0890/12, RGM 784 005/1, Seyithasan ( Turkey). RGM 784 005, Seyithasan ( Turkey), Fig. 15E .

Type locality. Seyithasan ( Turkey) .

Type stratum. Silt and clay of the Týrtar Formation.

Age. Middle Miocene, Serravallian.

Etymology. Referring to Anatolia in Turkey.

Diagnosis. Small, ovate-biconic shell with sculpture of broad, wide-spaced axial ribs overrun by prominent spiral cords, weakly shouldered last whorl with deep concavity between first two spiral cords, narrow elongate aperture with parietal and anal denticles almost touching, and relatively straight outer lip.

Description. Small, ovate-biconic shell of up to six teleoconch whorls; apical angle ~55°. Protoconch conical of 2.25 moderately convex whorls; diameter: 520 μm, height: 280 μm. Early teleoconch whorls almost flat sided with wide-spaced axial ribs, separated by wider interspaces, overrun by three prominent spiral cords. Later teleoconch whorls with weak mid-whorl angulation and prominent concavity between adapical and mid spiral cord. Last whorl high, attaining 70% of total height, weakly shouldered, moderately convex at periphery, moderately constricted at base, bearing nine wide-spaced axial ribs, fading over base, overrun by ten primary spiral cords over whorl and base, 1–2 delicate secondary spiral threads intercalated between primaries; fasciole slightly swollen, with prominent spiral cords, forming distinct umbilical chink. Aperture narrow, elongate. Columellar callus forming broad rim, sharply delimited from base. Columella moderately excavated in upper half, weakly twisted at siphonal canal. Anal canal narrowly U-shaped, deeply incised into thick alate adapical tip of aperture. Prominent parietal denticle and larger, broader anal denticle almost touching. Outer lip thickened by terminal varix, with crenulated margin and up to six prominent denticles close behind peristome; distance between anal denticle and adapical denticle slightly wider than that between lower denticles; denticles only slightly weakening abapically. Siphonal canal moderately long, moderately wide, slightly deflected to the left.

Discussion. Aplus anatolicus nov. sp. differs from the Paratethyan Aplus zebus ( De Gregorio, 1885 a) in its slenderer outline less excavated columella, and relatively straight outer lip. In addition, it has only one or two secondary spiral cord intercalated instead of three. Aplus exsculptus ( Dujardin, 1837) , from the Langhian of the Loire Basin, is morphologically closer to Aplus anatolicus compared to Aplus zebus , but differs in its squatter shape, tubercular sculpture and especially in its protoconch of only 1.5 whorls (see Glibert 1952: pl. 8, fig. 12 and Lozouet (2021: pl. 41, fig. 10, text-figs 6j–k).

Proto-Mediterranean Sea. Serravallian (Middle Miocene): Karaman Basin: Akboğaz at Lale River, Lale, Seyithasan ( Turkey) ( Landau et al. 2013).

Genus Dianthiphos Watters, 2009

Type species. Pisania bernardoi Costa & Gomes, 1998; original designation by Watters (2009: 257). Present-day, western Atlantic.

Original diagnosis. “ Fusiform; spire ca. 50% of length. Protoconch bulbous, 1.5 whorls, smooth, pink in the two known species. Teleoconch of 5 whorls, with spiral threads and axial ribs that become obsolete on last whorl. Single, thick, terminal varix. Columella angled at siphonal canal with a single denticle bounding anal canal. No internal lirae. Siphonal canal short, open.” ( Watters 2009: 257).

Included Miocene species. Hilda sacyi ( Cossmann & Peyrot, 1923) from the Burdigalian (Early Miocene) of Saucats ( France) (see Cossmann & Peyrot 1923: 297, pl. 16, fig. 46, pl. 17, figs 5–6, 19); Tritonidea productocostata ( Sacco, 1904) from the Burdigalian (Early Miocene) of the Colli Torinesi ( Italy) (see Sacco 1904: 59, pl. 14, figs 69–70); Pollia varians sensu Bellardi, 1873 [non Monostiolum varians ( Michelotti, 1847) ], from the Burdigalian and Langhian of the Colli Torinesi ( Italy) (see Bellardi 1873: 180, pl. 12, fig. 19); Pisania klaudiae Kovács & Vicián, 2023 , from the Badenian (Middle Miocene) of Hungary, Austria and Romania; Fusus philippi Michelotti, 1847 from the Tortonian (Late Miocene) of Stazzano ( Italy) (see Michelotti 1847: 277, pl. 9, fig. 20; Bellardi 1873: 179, pl. 12, fig. 18).

Stratigraphic and geographic range. Early to Late Miocene, (Burdigalian to Tortonian); Proto-Mediterranean Sea, Central Paratethys Sea, northeastern Atlantic, Present-day, western Atlantic.

Discussion. The genus was known so far only from two extant species from the western Atlantic: Dianthiphos bernardoi ( Costa & Gomes, 1998) and D. electrum Watters, 2009 . ‘Pisania’ klaudiae Kovács & Vicián, 2023 is especially similar to Dianthiphos bernardoi in shape and sculpture. It is difficult to define features that would reliably separate the Miocene species from Dianthiphos . Consequently, we suspect that Dianthiphos is rooted in the Miocene of the Circum-Mediterranean Region. Hilda Hoernes & Auinger, 1884 is distinguished from Dianthiphos by its stronger spiral sculpture, which is retained throughout ontogeny. Both genera seem to be closely related.

Figures

FIGURE 1. Measurements for shell morphology; AA: apical angle, SL: shell length, MD: maximum diameter, LWH: last whorl height, AH: aperture height, AL: aperture length, AW: aperture width.

FIGURE 2. Colubraria subobscura (Hoernes & Auinger, 1884). A1–A3. NHMW 1854/0035/0170, paralectotype, Lăpugiu de Sus (Romania). B1–B2. NHMW 1859/0037/0018a, paralectotype, Lăpugiu de Sus (Romania). C1–C3. NHMW 1865/0001/0183, paralectotype, Lăpugiu de Sus (Romania). D1–D2. NHMW 1859/0037/0018b, paralectotype, Lăpugiu de Sus (Romania). E1–E3. NHMW 1874/0025/0021, lectotype, Lăpugiu de Sus (Romania). F1–F2. NHMW 2023/0338/0001, paralectotype, Lăpugiu de Sus (Romania).

FIGURE 3. A. Colubraria subobscura (Hoernes & Auinger, 1884), NHMW 2023/0338/0002, Lăpugiu de Sus (Romania). B. Metula kovacsi nov. sp., NHMW 1867/0019/0078, holotype, CoŞteiu de Sus (Romania). C1–C2. Metula aliceae nov. sp. NHMW 1846/0037/0277a, paratype, Baden (Austria).

FIGURE 15. Protoconchs of Paratethyan Pisaniidae, Prodotiidae and Tudiclidae. A. Aplus moravicus (Hoernes & Auinger, 1890), NHMW 1862/0029/0034, lectotype, Rudice u Blanska (Czech Republic). B. Aplus subpusillus (Hoernes & Auinger, 1890), NHMW 1866/0001/1066, paralectotype, Forchtenau (Austria). C. Aplus transsylvanicus (Hoernes & Auinger, 1890), NHMW 1873/0026/0019b, lectotype. D. Aplus zebus (De Gregorio, 1885a), NHMW 1855/0045/0439, Gainfarn (Austria). E. Aplus anatolicus nov. sp. RGM 784 005, paratype, Seyithasan (Turkey). F. Hilda transsylvanica (Hoernes & Auinger, 1884), NHMW 1876/0011/0023a, lectotype, Lăpugiu de Sus (Romania).G. Janiopsis vindobonensis nov. sp., NHMW 1850/0009/0002, holotype, Baden (Austria). H. Monostiolum hungaricum (Csepreghy-Meznerics, 1954), NHMW 1868/0001/0094, Steinebrunn (Austria). I. Dianthiphos klaudiae (Kovács & Vicián, 2023), NHMW1846/0037/0144a, Steinebrunn (Austria). J. Euthria obelixi nov. sp., NHMW 1865/0015/0060, holotype, Lysice (Czech Republic). K. Tethyspollia mariae (Hoernes & Auinger, 1890), NHMW 1868/0001/0441, paralectotype, Lăpugiu de Sus (Romania).

FIGURE 21. Aplus anatolicus nov. sp. A1–A2. NHMW 1847/0058/0889, holotype, Seyithasan (Turkey). B1–B3. NHMW 1847/0058/0979, paratype Seyithasan (Turkey).