Aplus moravicus (Hoernes & Auinger, 1890)

Harzhauser, Mathias & Landau, Bernard M., 2024, The Colubrariidae, Eosiphonidae, Melongenidae, Pisaniidae, Prodotiidae and Tudiclidae (Gastropoda, Buccinoidea) of the Miocene Paratethys Sea, Zootaxa 5427 (1), pp. 1-110 : 30-31

Aplus moravicus (Hoernes & Auinger, 1890)

Figs 14A–C , 15A

* Pollia Moravica nov. form.—Hoernes & Auinger 1890: 238, pl. 28, fig 4a–b.

Type material. Lectotype (designated herein): NHMW 1862 /0029/0034, SL: 18.6 mm, MD: 10.6 mm, Rudice u Blanska ( Czech Republic), illustrated in Hoernes & Auinger (1890: pl. 28, fig. 4), Figs 14A 1 –A 2 , 15A . Paralectotypes: NHMW 2023 /0353/0001, SL: 19.4 mm , MD: 10.2 mm, Rudice u Blanska ( Czech Republic), Figs 14B 1 –B 2 . NHMW 2023 /0353/0002, SL: 15.5 mm , MD: 8.4 mm, Rudice u Blanska ( Czech Republic), Fig. 14C .

Revised description. Small, squat ovate-fusiform shell of up to 4.5 teleoconch whorls; apical angle 68–72°. Protoconch, broad conical of three moderately convex whorls; diameter: 900μm, height: 800μm. Last protoconch whorl with spiral cord placed just above abapical suture. First teleoconch whorl moderately convex with widely spaced axial ribs, overrun by three prominent spiral cords, strongly swollen over intersections; fourth weaker cord appears over subsutural ramp on second teleoconch whorl. Later teleoconch whorls convex, periphery slightly below mid-whorl. Three raised, narrow spiral cords appear on subsutural ramp on second teleoconch whorl. Suture strongly undulating around swollen intersections of abapical cord on preceding whorl. Interspaces between primary ribs deep, with 1–3 secondary spiral threads intercalated. Last whorl 70% of total height, with regularly convex periphery and strongly constricted base; sculpture of about 15 prosocline axial ribs overrun by about 12 strongly raised, convex cords, swollen over intersections; adapical three cords more closely spaced; fasciole broad, indistinct bearing strong spiral cords deeply cut by growth lines forming slightly scabrous surface. Aperture wide. Columella deeply excavated in upper half, angled at transition to siphonal canal, accentuated by prominent denticle at angulation. Columellar callus forming thin rim, moderately delimited from base. Columella with subobsolete denticles. Parietal denticle weak. Anal canal weakly incised, broadly U-shaped. No anal denticle. Outer lip weakly thickened with several narrow prominent lirae extending deep within, starting distinctly behind crenulated peristome. Siphonal canal long, very wide, slightly deflected to the left.

Discussion. Only subadult specimens are available and therefore, the parietal and anal denticles are not fully developed. Aplus moravicus (Hoernes &Auinger, 1890) is characterized by its prominent, regular, nodulose sculpture and evenly convex whorls. It is listed as synonym of the extant Mediterranean Aplus dorbignyi ( Payraudeau, 1826) by MolluscaBase Eds. (2023d). However, the two have little in common and Aplus dorbignyi is slenderer, has angled whorls with a broad subsutural ramp, coarser and less uniform tubercles and much more prominent denticles within the outer lip (see Aissaoui et al. 2016: figs 3A–B). Aplus multicostatus ( Bellardi, 1873) , from the Burdigalian or Langhian of the Colli Torinesi ( Italy), is morphologically much closer but differs in its higher last whorl, broader and more prominent axial ribs on the last whorl, and the prominent anal denticle (see Bellardi 1873: pl. 12, fig. 15; Ferrero Mortara et al. 1981: pl. 4, fig. 10).

Paleoenvironment. Shallow marine, inner neritic.

Distribution in Central Paratethys. Badenian (Middle Miocene): North-Alpine-Carpathian Foreland Basin: Rudice u Blanska ( Czech Republic) (Hoernes & Auinger 1890).

Figures

FIGURE 1. Measurements for shell morphology; AA: apical angle, SL: shell length, MD: maximum diameter, LWH: last whorl height, AH: aperture height, AL: aperture length, AW: aperture width.

FIGURE 2. Colubraria subobscura (Hoernes & Auinger, 1884). A1–A3. NHMW 1854/0035/0170, paralectotype, Lăpugiu de Sus (Romania). B1–B2. NHMW 1859/0037/0018a, paralectotype, Lăpugiu de Sus (Romania). C1–C3. NHMW 1865/0001/0183, paralectotype, Lăpugiu de Sus (Romania). D1–D2. NHMW 1859/0037/0018b, paralectotype, Lăpugiu de Sus (Romania). E1–E3. NHMW 1874/0025/0021, lectotype, Lăpugiu de Sus (Romania). F1–F2. NHMW 2023/0338/0001, paralectotype, Lăpugiu de Sus (Romania).

FIGURE 14. Aplus moravicus (Hoernes &Auinger, 1890). A1–A2.NHMW 1862/0029/0034, lectotype, Rudice u Blanska (Czech Republic). B1–B2. NHMW 2023/0353/0001, paralectotype, Rudice u Blanska (Czech Republic). C. NHMW 2023/0353/0002, paralectotype, Rudice u Blanska (Czech Republic).

FIGURE 15. Protoconchs of Paratethyan Pisaniidae, Prodotiidae and Tudiclidae. A. Aplus moravicus (Hoernes & Auinger, 1890), NHMW 1862/0029/0034, lectotype, Rudice u Blanska (Czech Republic). B. Aplus subpusillus (Hoernes & Auinger, 1890), NHMW 1866/0001/1066, paralectotype, Forchtenau (Austria). C. Aplus transsylvanicus (Hoernes & Auinger, 1890), NHMW 1873/0026/0019b, lectotype. D. Aplus zebus (De Gregorio, 1885a), NHMW 1855/0045/0439, Gainfarn (Austria). E. Aplus anatolicus nov. sp. RGM 784 005, paratype, Seyithasan (Turkey). F. Hilda transsylvanica (Hoernes & Auinger, 1884), NHMW 1876/0011/0023a, lectotype, Lăpugiu de Sus (Romania).G. Janiopsis vindobonensis nov. sp., NHMW 1850/0009/0002, holotype, Baden (Austria). H. Monostiolum hungaricum (Csepreghy-Meznerics, 1954), NHMW 1868/0001/0094, Steinebrunn (Austria). I. Dianthiphos klaudiae (Kovács & Vicián, 2023), NHMW1846/0037/0144a, Steinebrunn (Austria). J. Euthria obelixi nov. sp., NHMW 1865/0015/0060, holotype, Lysice (Czech Republic). K. Tethyspollia mariae (Hoernes & Auinger, 1890), NHMW 1868/0001/0441, paralectotype, Lăpugiu de Sus (Romania).