Aplus lapugyensis (Hoernes & Auinger, 1890)

Harzhauser, Mathias & Landau, Bernard M., 2024, The Colubrariidae, Eosiphonidae, Melongenidae, Pisaniidae, Prodotiidae and Tudiclidae (Gastropoda, Buccinoidea) of the Miocene Paratethys Sea, Zootaxa 5427 (1), pp. 1-110 : 28-29

Aplus lapugyensis (Hoernes & Auinger, 1890)

Figs 13A 1 –A 2

* Pollia Lapugyensis nov. form.—Hoernes & Auinger 1890: 239, pl. 28, figs 5–8.

Pollia lapugyensis Hö. Au. — Boettger 1902: 34.

Aplus lapugyensis (Hoernes et Auinger, 1890) — Kovács 2022: 88, figs 76–79.

Aplus lapugyensis (Hoernes et Auinger, 1885) — Kovács & Vicián 2023: 250, fig. 12C.

non Cantharus (Pollia) lapugyensis H. et Au. var.— Strausz, 1954: 26, pl. 4, figs 96a–b [= Aplus minutulus ( Bałuk, 1995) ]. non Cantharus (Pollia) lapugyensis (Hoern. & Auing.) — Báldi 1960: 67, pl. 2, fig. 7 [= Aplus subpusillus (Hoernes & Auinger, 1890) ].

Type material. Lectotype (designated herein), NHMW 1867 /0019/0082, SL: 22.6 mm, MD: 11.1 mm, CoŞteiu de Sus ( Romania), illustrated in Hoernes & Auinger 1890: pl. 28, fig. 5), Figs 13A 1 –A 2 . The other specimens illustrated by Hoernes & Auinger (1890: pl. 28, figs 6–8) seem to be lost.

Revised description. Medium-sized, moderately broad fusiform shell of about six teleoconch whorls; apical angle ~47°. Protoconch unknown. Early teleoconch whorls almost flat sided with periphery close above weakly incised suture. Sculpture of moderately prominent, close-set, orthocline axial ribs overrun by four spiral cords swollen at intersections. One or two weak secondary cords intercalated between primaries on third teleoconch whorl. Whorl profile becoming evenly convex on fourth whorl. Subsutural spiral cord becoming bifid on fourth whorl, accentuated by slight concavity below, especially on last whorl. Last whorl attaining 67% of total height, with convex periphery, constricted base, sculpture of about 11 axial ribs, most prominent along periphery, subobsolete over base, spiral sculpture of numerous weak primary and secondary cords swollen over axials. Fasciole broad, slightly swollen, forming narrow umbilical chink, bearing few spiral cords and marked growth lines. Aperture moderately narrow, ovate. Columella broadly excavated with six denticles variably developed, weakening adapically. Lowermost denticle coinciding with angled transition to siphonal canal. Anal canal moderately incised U-shaped, accentuated by narrow but prominent parietal denticle and larger anal denticle. Outer lip with weak terminal varix and eight, prominent, elongated denticles close behind peristome. Siphonal canal moderately short, wide, slightly deflected to the left, deeply notched.

Discussion. Aplus lapugyensis (Hoernes & Auinger, 1890) was regarded as a junior synonym of A. dorbignyi ( Payraudeau, 1826) in the literature (e.g., Brunetti & Della Bella 2014), however, the latter is a Pleistocene-Recent species, and is distinguished by its much stronger sculpture. Moreover, Aplus lapugyensis has more numerous axials on the early teleoconch whorls, and there are only three cords on the first whorl of Aplus dorbignyi , as opposed to four in Aplus lapugyensis . This species is reminiscent of Aplus hofae nov. sp. but has more delicate sculpture on the spire whorls and lacks the characteristic aspect of the primary cords that are divided into triplets Aplus hofae .

Paleoenvironment. Unknown; occurrences at Letkés ( Hungary) might point to inner neritic environments with corals ( Kovács & Vicián 2014).

Distribution in Central Paratethys. Badenian (Middle Miocene): Pannonian Basin: Letkés ( Hungary) ( Kovács & Vicián 2023); Făget Basin: Lăpugiu de Sus, CoŞteiu de Sus ( Romania) (Hoernes & Auinger 1890).

Figures

FIGURE 1. Measurements for shell morphology; AA: apical angle, SL: shell length, MD: maximum diameter, LWH: last whorl height, AH: aperture height, AL: aperture length, AW: aperture width.

FIGURE 2. Colubraria subobscura (Hoernes & Auinger, 1884). A1–A3. NHMW 1854/0035/0170, paralectotype, Lăpugiu de Sus (Romania). B1–B2. NHMW 1859/0037/0018a, paralectotype, Lăpugiu de Sus (Romania). C1–C3. NHMW 1865/0001/0183, paralectotype, Lăpugiu de Sus (Romania). D1–D2. NHMW 1859/0037/0018b, paralectotype, Lăpugiu de Sus (Romania). E1–E3. NHMW 1874/0025/0021, lectotype, Lăpugiu de Sus (Romania). F1–F2. NHMW 2023/0338/0001, paralectotype, Lăpugiu de Sus (Romania).

FIGURE 13. A1–A2. Aplus lapugyensis (Hoernes & Auinger, 1890), NHMW 1867/0019/0082, lectotype, CoŞteiu de Sus (Romania). B. Aplus minutulus (Bałuk, 1995), holotype, Z.PAL.U.W., No. BkK–G66S, Korytnica (Poland). C1–C2. Aplus hofae nov. sp., holotype, NHMW 1857/0024/0016, Lăpugiu de Sus (Romania).