Platystethus cornutus ( Gravenhorst, 1802 )

Lü, Liang & Zhou, Hong-Zhang, 2015, Review of the Genus Platystethus Mannerheim (Coleoptera: Staphylinidae: Oxytelinae) in China, Zootaxa 3915 (2), pp. 151-205 : 162-166

publication ID	https://doi.org/10.11646/zootaxa.3915.2.1
publication LSID	lsid:zoobank.org:pub:BEAD42C0-1031-4C82-A71A-15A670C3467A
DOI	https://doi.org/10.5281/zenodo.6114334
persistent identifier	https://treatment.plazi.org/id/03CE8795-FFD9-A449-FF11-A2B0FD52F871
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scientific name	Platystethus cornutus ( Gravenhorst, 1802 )
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3. Platystethus cornutus ( Gravenhorst, 1802) View in CoL

( Figs. 5–9 View FIGURE 5 View FIGURE 6 View FIGURE 7 View FIGURE 8 View FIGURE 9 )

Gravenhorst, 1802: 109 ( Oxytelus ; Type locality: Not cited, presumably Brunsvigae ( Herman, 2001a: 1476)); Mannerheim, 1830: 46 ( Platystethus ; Russia); Ganglbauer, 1895: 631 ( Platystethus ; Palaearctic region); Bernhauer & Schubert, 1911: 123 ( Platystethus ); Scheerpeltz, 1933: 1107 ( Platystethus ); Aleksandrov, 1934: 150 (Platysethus; China); Tottenham, 1939: 225 (type species of Craetopycrus ); Herman, 1970: 402 ( Platystethus ); Zheng, 1998: 255 ( Platystethus ; China); Herman, 2001a: 1476 ( Platystethus ); Herman, 2002: 100 ( Oxytelus ; senior homonym with Oxytelus cornutus Bernhauer, 1936 ); ICZN, 2003: 238 ( Oxytelus ; senior homonym); Smetana, 2004: 518 ( Platystethus , subgenus Craetopycrus ); Yan, Li & Zheng, 2012: 342 ( Platystethus ; China).

Synonymy.

Platystethus scybalarius Runde, 1835: 19 View in CoL (Type locality: Halae); Erichson, 1839: 586 (synonym of cornutus ).

Platystethus maxillosus Peyron, 1858: 432 View in CoL (Type locality: Not cited, presumably Tarsous ( Herman, 2001a: 1478)); Fauvel, 1872: 181 [= 1873: 23] (synonym of cornutus ).

Platystethus operosus Sharp, 1874: 91 View in CoL (Type locality: Japan); Fauvel, 1904: 46 (synonym of cornutus ); Bernhauer & Schubert, 1911: 124 (valid species); Bernhauer, 1922: 223 ( Formosa); Herman, 1970: 402 (valid species); Zheng, 1998: 255 (valid species; China); Herman, 2001a: 1485 (valid species); Smetana, 2004: 519 (subgenus Craetopycrus ; valid species); Yan, Li & Zheng, 2012: 342 (valid species; China).

Platystethus cornutus View in CoL race pallidipennis Bernhauer, 1906: 124 View in CoL (preoccupied by Staphylinus pallidipennis Panzer, 1795 View in CoL ; Type locality: transkaspisches Gebiet); Bernhauer & Schubert, 1911: 123 (variety of cornutus View in CoL ); Scheerpeltz, 1955: 83 (aberration of cornutus View in CoL ); Herman, 2001a: 1478 (synonym of cornutus View in CoL ); Smetana, 2004: 518 (subgenus Craetopycrus ; synonym of cornutus View in CoL ; homonym).

Platystethus cornutus View in CoL ab. nigripennis Gridelli, 1934: 72 (cited from Valle Indo: Skardu; Cachemir: Srinagar); Smetana, 1968: 236 (form of cornutus View in CoL ; Mongolia); Herman, 2001a: 1478 (Nomen nudum).

Platystethus cornutus View in CoL ssp. flavipennis Zheng, 2000: 112 View in CoL (preoccupied by Platystethus flavipennis Luze 1904 View in CoL ; Type locality: Xinjiang, Turpan [=Tulufan, 42.95ºN 89.19ºE], 30.VII.1995); Smetana, 2004: 518 (subgenus Craetopycrus ; subspecies of cornutus View in CoL ; homonym); Zheng, 2004: 310 (subgenus Craetopycrus ).

Platystethus cornutus View in CoL ssp. umbripennis Zheng, 2000: 112 (Type locality: Sichuan, Wolong [31.03ºN 103.18ºE], 1830m, 14.VII.1983); Smetana, 2004: 518 (subgenus Craetopycrus ; subspecies of cornutus View in CoL ); Yan, Li & Zheng, 2012: 342 (subspecies of cornutus View in CoL ).

Platystethus cornutus View in CoL ssp. stramineus Zheng, 2004: 310 (replacement name for ssp. flavipennis Zheng, 2000 View in CoL ); Yan, Li & Zheng, 2012: 342 (subspecies of cornutus View in CoL ).

Type material examined. [race pallidipennis Bernhauer ]. Syntype ( Fig. 7 View FIGURE 7 M): 1♀, Co-Type// Transcaspien K. Aris// v. pallidipennis Brh. Cotypus // Chicago NHMus M.Bernhauer Collection// Brit. Mus. 1956-709 ( BMNH).

[ssp. stramineus Zheng]. Paratypes ( Fig. 7 View FIGURE 7 A–F): 1♂, 1♀, Xinjiang, Tulufan [=Turpan, 42.95ºN 89.19ºE], Nanchong Teacher College [former name of CWNU, not locality but depository]// 1995.7.30, Collectors: H.-Q. Ji, L. Tan ( CWNU). [Note: all the CWNU specimens loaned from Prof. F.-K. Zheng were wrapped in ethanol cotton and preserved with labels in small glass vials sealed with adhesive tape. The identification and nature of specimens were written on the stopper and on the tape of each vial by Prof. Zheng.]

[ssp. umbripennis Zheng]. Paratypes ( Fig. 7 View FIGURE 7 G–L): 1♂, 1♀, Sichuan, Zigong [29.34ºN 104.78ºE], Nanchong Teacher College [former name of CWNU, not locality but depository]// 1982.VIII, Collectors: [not mentioned in label] ( CWNU).

[ operosus Sharp ]. Lectotype [designated here, Fig. 9 View FIGURE 9 ]: ♂, Type // Japan. G. Lewis.// Sharp Coll 1905-313.// Platysthetus [sic] operosus type D. S. ( BMNH).

Other material examined. 271 specimens, 125♂♂, 146♀♀. CHINA: 1♀, 1927. VI.29, E. Licent leg. (IZ-CAS); 1♀, 1914. V.28, E. Licent leg. [presumable], Platystethus cornutus ssp. operosus Sharp det. P.M.Hammond, 1980 (IZ-CAS); 1♂, same data as previous except: 1914. VI.16 (IZ-CAS). Beijing: 1♀, PEI-PING [=Beijing], 1942.X (IZ-CAS); 1♀, Beijing Botanical Garden [40.00ºN 116.21ºE], heap trap (weed), 2011. IX.3, D.-K. Zhou leg. ( IZ- CAS); 1♀, Chaoyang Dt., Datun Rd., IZ-CAS [40.00ºN 116.37ºE], 50 m, 2013. IV.24, F.- Y. Wang leg. (IZ-CAS); 31♂♂, 32♀♀, Chaoyang Dt., Datun Township [40.00ºN 116.41ºE], 1996. IV.21, H.-Z. Zhou leg. (IZ-CAS); 1♀, Changping, Er Bo Zi village [40.08ºN 116.31ºE], 70m, 1994. IV.17, H.-Z. Zhou leg. (IZ-CAS); 1♂, Haidian, Lan Dian Chang [39.96ºN 116.29ºE], 1997. V.16, H.-S. Zhou leg. (IZ-CAS); 1♀, Haidian, Yuan Ming Yuan [=Old Summer Palace, 40.00ºN 116.30ºE], 55m, 1993. VIII.2, H.-Z. Zhou leg. (IZ-CAS); 1♂, same data as previous except: 2004. III.25, H.-L. Shi leg. (IZ-CAS); 1♂, 6♀♀, Wo Fuo Si [=Reclining Buddha Temple, 40.00ºN 116.21ºE], 1980. IX.5 (IZ-CAS); 1♀, same data as previous except: 1980. IX.12, P.- Y. Yu leg. (IZ-CAS). Gansu: 1♀, Lanzhou, 1955. IV.29, S.-J. Ma, K.-L. Xia & Y.-L. Chen legg. (IZ-CAS); 2♂♂, same data as previous except: 1955. IV.30 (IZ-CAS). Hebei: 5♂♂, 2♀♀, Prov. Hopeh [=Hebei], Sienhsien [=Xianxian or Xian Co., 38.19ºN 116.12ºE], Platystethus cornutus ssp. operosus Sharp det. P.M.Hammond, 1980 (IZ-CAS); 1♂, HO-PEI [=Hebei], Sienhsien, 1937. IV.18, Platystethus cornutus ssp. operosus Sharp det. P.M.Hammond, 1980 (IZ-CAS); 1♂, Sienhsien, 1936. V.11 (IZ-CAS); 2♀♀, same data as previous except: 1936. VII.5 (IZ-CAS); 1♂, same data as previous except: 1938. VI.23 (IZ-CAS). Heilongjiang: 1♂, Harbin, 1945. V.18, Platystethus cornutus Grav. det. P.M.Hammond 1980 (IZ-CAS). Hubei, Shenlongjia, 1♂, Banqiao [31.41ºN 110.17ºE], 1998. VIII.5, H.-Z. Zhou leg. (IZ-CAS); 1♂, Xiagu [31.37ºN 110.24ºE], Zhaqigou, 600m, sweeping, 1998. VIII.6, C.-J. Ye leg. (IZ-CAS). Inner Mongolia: 1♂, 1♀, Ulanqab, Jining [41.00ºN 113.13ºE], 1972. VII.5, Platystethus cornutus Grav. det. P.M.Hammond 1980 (IZ-CAS). Jiangsu: 2♂♂, KIANGSU [=Jiangsu Prov.], “YU-TOAN”, Platystethus cornutus ssp. operosus Sharp det. P.M.Hammond, 1980 (IZ-CAS); 1♂, 1♀, Nordwestl. China [?], Chinkiang [=Zhenjiang?], Col. Reitter, operosus Shp. Det. Bernhauer, Chicago NHMus M. Bernhauer Collection ( FMNH). Jiangxi: 1♂, Kiu Kiang [=Jiujiang, 29.70ºN 116.00ºE], Sharp Coll, 1905-313, Platystethus cornutus var. collaris Brnh., Chicago NHMus M. Bernhauer Collection, Platystethus cornutus (Gr.) det. P.M.Hammond 1969 ( FMNH). Qinghai: 1♂, 3♀♀, Chaka Township [36.79ºN 99.08ºE], 3200 m, under rock, 2005. IX.10, J. Wu leg. (IZ-CAS); 1♀, Ge’ermu [=Golmud, 36.40ºN 94.91ºE], 2700m, 1961. IV.21, L.- Y. Wang leg., Platystethus cornutus Grav. det. P.M.Hammond 1981 (IZ-CAS). Shaanxi: 8♂♂, 9♀♀, Heyangqiao, Xingzhuang, 1997. V.30, H.-S. Zhou leg. ( IZ- CAS). Shandong: 2♀♀, Tsinanfou [=Jinan], Long-tong, 500 à 700 m, Platystethus cornutus ssp. operosus Sharp det. P.M.Hammond, 1980 (IZ-CAS); 1♂, Tsingtao [=Qingdao], Platystethus cornutus ssp. operosus Sharp det. P.M.Hammond, 1980 (IZ-CAS). Shanghai: 2♂♂, 1947. VIII.1, Platystethus cornutus ssp. operosus Sharp det. P.M.Hammond, 1980 (IZ-CAS); 32♂♂, 43♀♀, Prov. KIANGSU, Shanghai, Zi-ka-wei [=Xujiahui], 1939. III.20–1940, Platystethus cornutus ssp. operosus Sharp det. P.M.Hammond, 1980 (IZ-CAS); 9♂♂, 13♀♀, “? Zi-ka-wei” (IZ-CAS). Sichuan: 2♂♂, 2♀♀, Baoxing, Longdong [30.47ºN 102.71ºE], Ruobigou, 2003. VIII.13, grass heap by road, X.-D. Yu leg. (IZ-CAS). Tientsin [= Tianjin]: 4♂♂, 5♀♀, 1938.IV, Platystethus cornutus ssp. operosus Sharp det. P.M.Hammond, 1980 (IZ-CAS). Xinjiang: Bachu [=Maralbexi], 1350 m, 1958. IX.18, C.-Q. Li leg., Platystethus cornutus ab. pallidipennis Bnh. det. P.M.Hammond 1980 (IZ-CAS); 2♂♂, 5♀♀, Kashi [=Kashgar, 39.45ºN 76.01ºE], 1959. VI.17, from pot ale, “5417 4” [meaning unknown], Platystethus cornutus ssp. operosus Bnh. det. P.M.Hammond 1980 (IZ-CAS); 1♂, 1♀, Ku’erle [= Koria or Korla, 41.73ºN 86.17ºE], 2008. VI.1, H. Huan leg.(IZ-CAS); 1♂, Ruoqiang [=Qarkilik, 39.02ºN 88.16ºE], Alagan, 850 m, 1960. V.7, S.- Y. Wang leg., Platystethus cornutus var. pallidipenne [sic] Brnh. det. P.M.Hammond 1981 (IZ-CAS); 2♂♂, Shanshanxian [=Shanshan Co. or Piqan Co. in Turpan Prefecture, 42.87ºN 90.21ºE], 22–350 m, 1958. VI.22, C.-Q. Li leg., Platystethus cornutus ab. pallidipennis Bnh. det. P.M.Hammond 1980 (IZ-CAS); 2♂♂, Shufu [=Konaxeher, 39.38ºN 75.86ºE], 1200 m, 1959. VI.20, A-Fu Tian leg., Platystethus cornutus var. pallidipenne [sic] Brnh. det. P.M.Hammond 1981 (IZ-CAS); 4♀♀, same data as previous except: Jiashi [=Peyziwat or Payzawat, 39.48ºN 76.72ºE], 1160 m (IZ-CAS); 1♀, Shule Co. [39.40ºN 76.05ºE], 1959. VI.13, oil well, 5365 4 [meaning unknown], Platystethus cornutus ssp. operosus Bnh. det. P.M.Hammond 1980 (IZ-CAS); 1♀, Tuokexun [=Toksun, 42.79ºN 88.65ºE], 1958. VI.17, G. Wang leg., Platystethus cornutus ab. pallidipennis Bnh. det. P.M.Hammond 1980 (IZ-CAS). Xizang: 1♀, Lhasa, Bayi Farm [29.83ºN 91.58ºE], 3660m, 1960. V.14, C.-G. Wang leg., Platystethus cornutus Grav. det. P.M.Hammond 1980 (IZ-CAS). BELGIQUE: 1♂, “Werner” [hand-writing not clear] ( IZ- CAS). FRANCE: 1♂, Le Bourget, 1935. VII.7, Platystethus arenarius Fourc. det. anonym (IZ-CAS). HUNGARY: 1♂, Siófok, F. Lichtneckert leg. [unreliable] ( HNHM); 1♂, 1♀, “Hu. c. Dömsöd” [a village in Pest county, Hungary], “dömsödi legelő” [=Dömsöd pasture], “dömsödi szikes legelő” [=Dömsöd saline pasture], 1953. VII.15, Z. Kaszab leg. C. cornutus (Grav.) , det. Adám 1983, ( HNHM); 1♀, “Hu. occ. Somogy-m. Zamárdi” [a town in Somogy county, Hungary], “Töreki láp, IX.8, IV tó, ripicol” [Töreki moor], 1953. IX.8, Dr. Kaszab leg. ( HNHM).

Redescription. Body black to dark brown to reddish brown; mandibles, maxillary palpi, antennae, and legs lighter; elytra ( Figs. 7 View FIGURE 7 A, B, G, H, M; 8B–E, H–J) lighter than head and pronotum, colored variously, dark brown to reddish brown to entirely pale, in some cases with various pale patterns. Length [average]: ♂, 3.3 mm; ♀, 3 mm.

Male. Head ( Figs. 5 View FIGURE 5 A; 6A) broadest at eyes or temples. Disc coriaceous, nearly glabrous, with punctures dense and coarse. Clypeus transverse and protruding beyond anterior border of supra-antennal ridges, with anterior margin truncate in middle, furnished a pair of sharp and slender spines. Epistomal suture with middle portion absent, lateral portions short but obvious and posteromedially directed. Supra-antennal ridges obviously elevated. Vertex depressed in anterior part, convex in center; mid-longitudinal suture short, posteriorly reaching occipital suture. Eyes slightly convex, shorter than temples, with fine facets; orbital sulcus feeble. Long and deep longitudinal groove sculptured throughout mesial side of eye and temple. Temple not dilated laterally. Neck not constricted. Occipital suture slightly curved, present only in middle, with large setal fovea and feeble paralateral suture lying at each end. Mandible ( Fig. 6 View FIGURE 6 C) long, stout, and incurved at apex; bearing 3 small teeth on inner edge; with tiny seta at middle of dorsal surface.

Pronotum ( Figs. 5 View FIGURE 5 A; 6A) slightly wider than head, widest at near anterior margin, with mid-longitudinal sulcus present whole length, with punctures sparser and smaller than in head, with micro-striae almost absent; anterior margin slightly bi-emarginate, anterior lateral angles produced, no posterior angles; lateral and posterior margins integrated and evenly rounded, without crenulation. Scutellar impression heart-shaped, anterior margins crescent. Elytra with posterior margin truncate, elytral suture dehiscent ( Fig. 7 View FIGURE 7 H) or partly overlapping ( Fig. 8 View FIGURE 8 B).

Abdomen coriaceous. Sternite VII ( Figs. 5 View FIGURE 5 C; 6E) with two tiny teeth on posterior margin, between which shallowly emarginate. Sternite VIII ( Figs. 5 View FIGURE 5 D; 6F) tri-partitioned by 2 curved narrow sutures, middle part truncate on posterior margin, without mid-longitudinal internal ridge, lateral parts separated by middle part but connected at anterior margin, each with two teeth at mesial side of posterior margin: one sharp, one dull; basal ridge broken in middle and subbasal ridge with middle portion absent.

Aedeagus ( Figs. 5 View FIGURE 5 G–I; 6I –L) with median lobe inflated at base and gradually narrowed apically; apical orifice fissured to middle of ventral surface, edges of ventral fissure sharply protruding into a pair of long spines; internal sac membranous and rolling into watch-spring-shape at base ( Figs. 5 View FIGURE 5 H; 6K; 7E, K). Paramere arm-like, basal arm with furrow on ventral surface; apical arm explanate and furnished with small seta at apex, one or two setae at near apical 1/3, and with pits scattered.

Female. Head and pronotum smaller ( Figs. 5 View FIGURE 5 B; 6B); clypeus more protruding, anterior margin truncate with two reduced denticles; mid-longitudinal suture nearly reduced; eyes more convex. Mandibles ( Fig. 6 View FIGURE 6 D) shorter than in male, with 3 teeth prominent. Pronotal anterior margin ( Figs. 5 View FIGURE 5 B; 6B) slightly bi-emarginate or nearly straight. Abdominal sternite VII with posterior margin straight. Sternite VIII ( Figs. 5 View FIGURE 5 E; 6G) not partitioned, without teeth, with posterior margin broadly rounded and a little protruding in middle.

Spermatheca ( Figs. 5 View FIGURE 5 F; 6H) comma-formed, basal part inflated into globoid, apical part thinner and shorter, a little curved, and sharp in apex.

Distribution. China (Beijing, Gansu, Hebei, Heilongjiang, Hong Kong, Inner Mongolia, Jiangsu, Jiangxi, Jilin, Liaoning, Ningxia, Qinghai, Shanghai, Sichuan, Shaanxi, Shandong, Taiwan, Tianjin, Xizang, Xinjiang); Europe, Georgia, Russia, Mongolia, Madeira, Algeria, Canary Islands, Morocco, Algeria, Libya, Sudan, Egypt, Saudi Arabia, Syria, Turkey, Iraq, Iran, “Turkestan”, Afghanistan, Cyprus, Israel, N. & S. Korea, Uzbekistan, Kazakhstan, Tajikistan, Turkmenistan, Pakistan, India, Nepal, Myanmar, Vietnam, Japan.

Remarks. Sharp (1874: 91) described the species P. operosus and differentiated it from P. cornutus by “smaller and with the head less strongly punctured”. The size of P. operosus in Sharp’s description is “1 ½ Lin.”, which is same as the measure in Gravenhorst (1802: 109). Sharp declared that there were four specimens, but he did not mention their genders. We examined the male syntype specimen ( Fig. 9 View FIGURE 9 , designated as lectotype in this paper) of P. operosus from Sharp’s collection (BMNH) and saw pictures of the other three syntypes (2♂♂1♀ with similar label data) in Lewis’ collection (BMNH). Comparing them with all the P. cornutus specimens we have from Europe and China, we conclude that the type of P. operosus does not differ from P. cornutus beyond the normal range of variation of the latter ( Figs. 6–8 View FIGURE 6 View FIGURE 7 View FIGURE 8 ). So we agree with Fauvel (1904) and re-synonymize P. operosus under P. cornutus . The following is a brief comment on the spines on aedeagal median lobe: at first sight, they look like parts of internal sac everted from the capsule of median lobe, but according to our observation, they are connected with the wall of median lobe by nearly transparent membrane. And possibly two “strips” of internal sac “touch” the inside base of the spines. Therefore they should be parts of capsule, which is logically consistent with the similar structure in other species like P. arenarius etc. The species in cornutus group have same structures and similar forms in aedeagus. Even the internal sac, normally species-specific in many situations, is hardly distinguishable. The paired spines on median lobe are variable (in lateral view) among species and stable within species. And the other useful treat is the diverse parameral apex.

References

Aleksandrov, A. I. (1934) The list of beetles of the family Staphilinidae (Coleoptera) of Girin province with description of new forms [K сpiсku zukov сiмiйсtva Staphilinidae (Coleoptera) iz Giрiнсkoй pрoviнtii с opiсaнiiм нovыk фoрм]. Yearbook Club of Natural History and Geography Young Men's Christian Association, Harbin [Izigodнik kлuba iсtiсtvozнaнii i giogрaфii Kрiсtiaнсkogo сoюza мoлodыk люdiй, Kaрbiн], 1 (1933), 150 - 155.

Bernhauer, M. (1906) Neue Staphyliniden der palaarktischen Fauna nebst synonymischen Bemerkungen. Munchener Koleopterologische Zeitschrift, 3, 123 - 128.

Bernhauer, M. & Schubert, K. (1911) Staphylinidae II (Pars 29). In: Junk, W. & Schenkling, S. (Eds.), Coleopterorum Catalogus. Vol. V. W. Junk, Berlin, pp. 87 - 190.

Bernhauer, M. (1922) Sauter's Formosa-Ausbeute: Stapliylinidae I. Archiv fur Naturgeschichte Berlin. Abteilung A, 88, 220 - 237.

Erichson, W. F. (1839) Die Kafer der Mark Brandenburg. Vol. 1 (2). F. H. Morin, Berlin, 356 pp. [pp. 385 - 740]

Fauvel, A. (1872) Faune Gallo-Rhenane ou species des insectes qui habitent la France, la Belgique, la Hollande, le Luxembourg, la prusse Rhenane, la Nassau et la Valais avec tableaux synoptiques et planches gravees. Vol. 3. Livraison 3. F. le Blanc-Hardel, Caen, 214 pp.

Fauvel, A. (1904) Staphylinides de l'Hindoustan et de la Birmanie. Revue d'Entomologie publiee par la Societe Francaise d'Entomolodie, 23, 43 - 70. [Caen]

Ganglbauer, L. (1895) Die Kafer von Mitteleuropa. Die Kafer der osterreichisch-ungarischen Monarchie, Deutschlands, der Schweiz, sowie des franzosischen und italienischen Alpengebietes. 2. Familienreihe Staphylinoidea. Theil I. Staphylinidae, Pselaphidae. Carl Gerold's Sohn, Wien, 881 pp.

Gravenhorst, J. L. C. (1802) Coleoptera Microptera Brunsvicensia nec non exoticorum quotquot exstant in collectionibus entomologorum Brunsvicensium in genera familias et species distribuit. Vol. 1. Carolus Reichard, Brunsuigae, lxvi + 206 pp.

Gridelli, E. (1934) Materiali zoologici raccolti dalla spedizione italiana al Karakoram (1929 - Anno VII). Coleoptera Staphylinidae. Atti del Museo Civico di Storia Naturale di Trieste, 12, 69 - 85.

Herman, L. H. (1970) Phylogeny and reclassification of the genera of the rove-beetle subfamily Oxytelinae of the world (Coleoptera, Staphylinidae). Bulletin of the American Museum of Natural History, 142, 345 - 454.

Herman, L. H. (2001 a) Catalog of the Staphylinidae (Insecta, Coleoptera): 1758 to the end of the second millennium. vol. 3, Oxyteline group. Bulletin of the American Museum of Natural History, No. 265, 1067 - 1806.

Herman, L. H. (2002) Case 3207. Staphylinidae Latreille, 1804 (Insecta, Coleoptera): Proposed conservation of 65 specific names. Bulletin of Zoological Nomenclature, 59, 99 - 113.

ICZN (2003) Opinion 2053 (Case 3207). Staphylinidae Latreille, 1804 (Insecta, Coleoptera): 65 specific names conserved. Bulletin of Zoological Nomenclature, 60, 237 - 246.

Luze, G. (1904) Beitrag zur Staphyliniden-Fauna von Russisch-Central-asien (Coleoptera). Horae Societatis Entomologicae Rossicae, 37, 74 - 115.

Mannerheim, C. G. von (1830) Precis d'un nouvel arrangement de la famille des brachelytres de l'ordre des insectes coleopteres. St. Petersbourg, 87 pp.

Panzer, G. W. F. (1795) Entomologia germanica exhibens insecta per germaniam indigena secundum classes, ordines, genera, species adiectis syonymis, locis, observationibus. I. Eleuterata. Felsecker, Normibergae, 8 + 12 + 370 + 2 pp.

Peyron, E. (1858) Catalogue des Coleopteres des environs de Tarsous (Caramanie), avec la description des especes nouvelles. Annales de la Societe Entomologique de France. Series 3, 6, 353 - 434.

Runde, G. H. (1835) Brachelytrorum species Agri Halensis. Ploetzianis, Halae, 6 + 32 pp.

Scheerpeltz, O. (1933) Staphylinidae VII (Pars 129). In: Junk, W. & Schenkling, S. (Eds.), Coleopterorum Catalogus. Vol. VI. W. Junk, Berlin, pp. 989 - 1500.

Scheerpeltz, O. (1955) Eine neue Art der Gattung Platystethus Mannh., mit einer Bestimmungstabelle der westpalaarktischen Arten und Formen dieser Gattung (Col. Staphylinidae). Koleopterologische Rundschau, Vienna, 33, 78 - 88.

Sharp, D. S. (1874) The Staphylinidae of Japan. Transactions of the Entomological Society of London, 1874, 1 - 103.

Smetana, A. (1968) Ergebnisse der zoologischen Forschungen von Dr. Z. Kaszab in der Mongolei. 127. Staphylinidae III. Unterfamilien Oxytelinae und Oxyporinae (Coleoptera). Acta entomologica Bohemoslovaca, 65, 226 - 238.

Smetana, A. (2004) Subfamily Oxytelinae Fleming, 1821. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera: Hydrophiloidea - Histeroidea - Staphylinoidea. Vol. 2. Apollo Books, Stenstrup, pp. 511 - 535.

Tottenham, C. E. (1939) Some notes on the nomenclature of the Staphylinidae (Coleoptera). Proceedings of the Royal Entomological Society of London (B), 8, 224 - 226, 227 - 237.

Yan, X. - H., Li, Y. - J. & Zheng, F. - K. (2012) A new species of the genus Platysyethus from China (Coleoptera, Staphylinidae, Oxytelinae). Acta Zootaxonomica Sinica, 37, 341 - 344.

Zheng, F. - K. (1998) A taxonomic study on Staphylinidae in China, Oxytelinae: Platystethus. Journal of Sichuan Teachers College (Natural Science), 19, 249 - 256.

Zheng, F. - K. (2000) Notes on Chinese species of the genus Platystethus Mannerheim (Coleoptera: Staphylinidae: Oxytelinae). In: Zhang, Y. - L. (Ed.), Systematic and faunistic research on Chinese insects. Proceedings of the 5 th National Congress of Insect Taxonomy. China Agriculture Press, Beijing, pp. 110 - 116.

Zheng, F. - K. (2004) A new name in the genus Platystethus Mannerheim (Coleoptera: Staphylinidae, Oxytelinae). Coleopterists Bulletin, 58, 310.

Figures

FIGURE 5. Platystethus cornutus (Gravenhorst, 1802) I: A, head and pronotum ♂ (dorsal view); B, head and pronotum ♀ (dorsal view); C, sternite VII ♂; D, sternite VIII ♂; E, sternite VIII ♀; F, spermatheca; G, aedeagus (ventral view); H, median lobe of aedeagus (lateral view); I, right paramere (in situ) of aedeagus (lateral view). Scales: A, B = 0.5 mm; F = 0.05 mm; rest, 0.3 mm.

FIGURE 6. Platystethus cornutus (Gravenhorst, 1802) II: A, head and pronotum ♂ (dorsal view); B, head and pronotum ♀ (dorsal view); C, right mandible ♂; D, right mandible ♀; E, sternite VII ♂; F, sternite VIII ♂; G, sternite VIII ♀; H, spermatheca; I, aedeagus (ventral view); J, aedeagus (dorsal view); K, median lobe of aedeagus (lateral view); L, right paramere (in situ) of aedeagus (lateral view). Scales: H = 0.05 mm; rest, 0.2 mm.

FIGURE 7. Platystethus cornutus (Gravenhorst, 1802) III: A – F, paratypes of ssp. stramineus Zheng, 2004; G – L, paratypes of ssp. umbripennis Zheng, 2000; M, syntype of “ race pallidipennis ” Bernhauer, 1906. A, G, forebody ♂ (dorsal view); B, H, M, forebody ♀ (dorsal view); C, I, aedeagus (ventral view); D, J, aedeagus (dorsal view); E, K, median lobe of aedeagus (lateral view); F, L, right paramere (in situ) of aedeagus (lateral view). Scales: 0.3 mm.

FIGURE 8. Platystethus cornutus (Gravenhorst, 1802) IV (specimens from different localities): A, France; B, Hungary (Dömsöd); C, Hungary (Siófok); D, Shaanxi; E, F, K, Beijing (K — aedeagus in ventral view); G, H, L, Shanghai (G — male, H — female, L — aedeagus in ventral view with internal sac partly everted); I, Jiangxi (Jiujiang); J, Jiangsu (Chinkiang [= Zhenjiang]). Scales: 0.3 mm.

FIGURE 9. Platystethus cornutus (Gravenhorst, 1802) V (Lectotype of P. operosus Sharp, 1874 in BMNH): A, habitus; B, forebody; C, sternite VII; D, sternite VIII; E, F, aedeagus (E — ventral view, F — dorso-lateral view); G – I, median lobe of aedeagus (G — lateral view, H — dorsal view, I, apical view); J, right paramere (in situ) of aedeagus (lateral view); K, specimen on mounting card with labels. Scales: A, B = 0.5 mm; rest, 0.2 mm.