Pinguicula macroceras Link (1820: 54)

Domínguez, Yoannis, Valdés, Cristina Mercedes Panfet & Miranda, Vitor Fernandes Oliveira, 2017, Typification of names in the genus Pinguicula L. (Lentibulariaceae), Phytotaxa 312 (2), pp. 179-198 : 187-188

Pinguicula macroceras Link (1820: 54)

Protologue locality:—Unalaschka.

Type (neotype, designated here):—[ UNITED STATES OF AMERICA]. Unalaschka, s.d., J. F. Eschscholtz s.n. ( LE s.n. [digital photo!], the five plants on the left side of the sheet under the name Pinguicula vulgaris L. var. macroceras Herd. ; Fig. 2a ).

≡ Pinguicula vulgaris Linnaeus (1753: 17) var. macroceras (Link) Herder (1872: 380) .

≡ Pinguicula vulgaris L. subsp. macroceras (Link) Calder & Taylor (1965: 1399) .

= Pinguicula breviscapa Ledebour ex Herder (1872: 381) , nom. nud.

= Pinguicula camtschatica Willdenow ex Link (1820: 54) , nom. nud.

= Pinguicula macroceras Link subsp. nortensis Steiger & Rondeau in Rondeau & Steiger (1997: 3). Type (holotype):—[ UNITED STATES OF AMERICA]. Del Norte Co., S. Fork Smith River, Hiouchi, at Sheep Pen Creek x Douglas Park Rd. (T16N-R1E-sw1/4 sw1/4 sec 10), 100 m, 7 May 1971, J. Steiger s.n. (BERN).

= Pinguicula microceras Chamisso (1831: 568) ≡ Pinguicula macroceras Link var. microceras (Cham.) Casper (1962: 116) . Type (lectotype, designated here):—[ UNITED STATES OF AMERICA]. Unalaschka, s.d., s. coll. [A. Chamisso] (LE s.n. [digital photo!], plants in the lower half of the sheet, isolectotype MO No. 1764211 barcode MO-149399 [digital photo!]; image of the isolectotype is available at http://www.tropicos.org/Image/34179).

= Pinguicula unalaccensis Fischer ex Herder (1872: 380) . Type (lectotype, designated here):—[ UNITED STATES OF AMERICA]. Unalaschka , s.d., Langsdorff s.n. (LE s.n. [digital photo!], the four plants on the right side of the sheet; Fig. 2b ).

= Pinguicula vulgaris L. f. albiflora Komiya (1972: 47) . Type (holotype):— JAPAN. Tohoku, Mt. Kurikoma, Miyagi, Iwate, 1890, S. Ikeno & T. Makino s.n. (TI s.n.).

= Pinguicula vulgaris L. var. floribunda Watanabe & Takeda in Takeda & Watanabe (1997: 237). Type (holotype):— JAPAN. Honshu, Mt. Sengoku, ca. 850 m alt., Okunohira-tani, Iitaka, Mie Prefecture, 5 June 1996, S. Watanabe & A. Takeda s.n. (TI s.n. [digital photo!]).

Note:— Pinguicula macroceras was described by Link (1820: 54) from a specimen collected by P.S. Pallas in Unalaska (Pallas 422, in Herb. Willd.).According to Sleumer (1949), Pilger (1953) and Hiepko (1987) the Willdenow’s herbarium (B-W) was totally saved from destruction during the bombing raid in Berlin on 1943; nonetheless, according to Casper (1962a, 1962b, 1966), the type specimen of P. macroceras was destroyed during the war. We have unsuccessfully searched for this specimen at B and among the records of materials destroyed at Berlin; and yet, no other original material (Pallas No. 422) was found in any collection searched by us. In Iconographia Botanica seu Plantae Criticae ( Reichenbach 1823: pl. 82, f. 169–170) appears a drawing of P.macroceras based on a specimen from B-W, but Chamisso (1831: 568) stated that it represented his own P. microceras , a taxon also from Unalaska which is currently considered a synonym of P. macroceras . We have found several specimens in different herbaria identified as P. macroceras but most of them are types for other names synonymized under P. macroceras and they do not represent Link’s name concept. In the herbarium of the V.L. Komarov Botanical Institute (LE) we have found two specimens from Unalaska which show the main character in Link’s diagnosis: “calcare turbinate corollae laciniis longiore”; therefore, they are suitable for the designation of a neotype. The first sheet is an admixture specimen containing a collection made by J.F. Eschscholtz in Unalaska and identified as “ Pinguicula vulgaris L. var. macroceras Herd. ”, which is a homotypic synonym of P. macroceras ( Herder 1872: 380) . The second sheet is under the name P. macroceras , but we could not identify the collector’s name or other annotations in the labels. Hence, we have selected as neotype the plants collected by Eschscholtz in the LE admixture specimen ( Fig. 2a ). In the same sheet appears a collection made by Langsdorff in Unalaska which corresponds to Herder’s name Pinguicula unnalascensis and is here designated as lectotype for this name ( Fig. 2b ).

Figures

FIGURE 2. Neotype of Pinguicula macroceras Link (a) and lectotype of Pinguicula unalaccensis Fisch. ex Herder (b), at the Herbarium of the V.L. Komarov Botanical Institute (LE). Photography by courtesy of the herbarium LE; reproduced with permission.