Lasianthus cyanocarpus Jack (1823:125)

Napiroon, Tiwtawat, Chayamarit, Kongkanda, Dawson, Sally, Till, Walter & Balslev, Henrik, 2020, A synopsis of Lasianthus (Lasiantheae, Rubiaceae) in Thailand and two additional new species, Phytotaxa 439 (1), pp. 1-38 : 14

15. Lasianthus cyanocarpus Jack (1823:125) . Type (neotype designated by Zhu 2002):― INDONESIA. Sumatra,

west coast hill, 15 June 1953, J van W. Borssum 1864 (neotype L0306104 image!). ( Figure 3G ).

= Lasianthus inaequalis Blume (1826–1827:996). Type (lectotype designated by Zhu 2012):― INDONESIA. Java, Seribu, C. L. Blume s.n. (lectotype L0000688 image!).

= Lasianthus cyanocarpus var. subsessilis King & Gamble (1904:113) . Type (lectotype designated here): — MALAYSIA. Goping, August 1880 (exact date not given), Dr. King’s Collector s.n. (lectotype K000763940!).

= Lasianthus bractescens Ridley (1918:85) . Type (lectotype designated here): — MALAYSIA. Selangor, Batu Caves woods, December 1896, H. N. Ridley 8231 (lectotype K000763941!).

T hailand: —PENINSULAR. Nakhon Si Thammarat: Khao Luang National Park, 1 April 2017, T. Napiroon & S. Vajrodaya 027 (BKF). Satun: Thaleban National Park, 3 November 1990, K. Larsen et al. 41087 (AAU, BKF), 30 January 2016, T. Napiroon & S. Vajrodaya 006 (BKF). Yala: 22 October 1970, C. Charoenphol, K. Larsen & E. Warncke 4151 (BKF, K).

Distribution: — Malaysia (Peninsular, Borneo), Indonesia (Sumatra, Java, Kalimantan, Lesser Sunda Islands, Irian Jaya), Philippines, Papua New Guinea, Thailand (Peninsular).

Ecology: —Evergreen forest and moist evergreen forest, in shaded places and near small streams at an elevation of 100‒ 900 m. Flowering in April‒June and fruiting in July‒September.

Notes: ―The holotype of Lasianthus cyanocarpus was destroyed by fire in 1824 and a neotype was designated ( Zhu 2012). Lasianthus cyanocarpus has been confused with L. hirsutus (Roxb.) Merr. (1952:229) because of their overall resemblance, especially in having leaf-like inflorescence bracts. In Thailand, a few species of Lasianthus have leaf-like inflorescences bracts, and in our key, we have grouped those species together. From our survey in the field and on herbarium specimens, L. cyanocarpus is restricted to peninsular Thailand and its characters agree with the protologue ( Jack, 1823).

Lasianthus cyanocarpus is closely related to L. hirsutus , but differs by having: relatively small leaves 8–13.5 × 2.5– 4.5 cm (not 13–20× 5–6 cm) that are cordate or slightly oblique at their bases; short petioles, 2‒3 mm (not 10–15 mm long); stipules inconspicuous, tomentose and persistent (not stipules conspicuous, densely hirsute and relatively permanent); bracteoles numerous and densely tomentose (not absent); drupes ovoid with 4–5 pyrenes (not globose with 5 pyrenes).

Figures

FIGURE 3A‒O. Lasianthus werefound in Thai naturalforests:A) L. appressus;B) &C) L.attenuatus;D) L.chinensis;E) L.chrysoneurus; F) L. constrictus; G) L. cyanocarpus; H) L. ellipsoideus; I) L. ellipticus; J) L. fordii; K) L. foxworthyanus; L) & M) L. hirsutus; N) L. hispidulus and O) L. inodorus.