Lasianthus hirsutus (Roxb.) Merr. (1952: 229)

Napiroon, Tiwtawat, Chayamarit, Kongkanda, Dawson, Sally, Till, Walter & Balslev, Henrik, 2020, A synopsis of Lasianthus (Lasiantheae, Rubiaceae) in Thailand and two additional new species, Phytotaxa 439 (1), pp. 1-38 : 17

24. Lasianthus hirsutus (Roxb.) Merr. (1952: 229) . ≡ Triosteum hirsutum Roxburgh (1824:180) . Type: ―S. loc., S.

coll. 8305B (holotype not seen; isotype A00053971 image!). ( Figure 3L & 3M ).

= Lasianthus oculuscati Miquel (1857:315) . Type: ― INDONESIA. Java, s. d., C. G. C. Reinwardt s.n. (holotype L0057762 image!; paratype L0057763 image!).

= Lasianthus cyanocarpus Jack var. novoguineensis Valeton (1911:498) . Type: ― INDONESIA. South-west New Guinea, s. d., G. M. Versteeg 1406 (holotype not seen; isotype L0310256 image!).

= Lasianthus cyanocarpus Jack var. asperatus Pierre ex Pitard (1924:381) . Type: ― VIETNAM. Bien Hoa, s. d., J. B. L. Pierre 3256 (syntype L0305709 image!).

= Lasianthus cyanocarpus Jack var. lucidulus Pierre ex Pitard (1924:382) . Type: ― VIETNAM. Bien Hoa, September 1869, J. B. L. Pierre, A. C. H. Alleizette s.n. (syntype L0310260 image!).

T hailand: —EASTERN. Surin: 2 December 1976, C. Phengklai 3527 (BKF). CENTRAL. Nakhon Nayok: Khao Yai National Park, 11 October 1999, S. Chongko 39 (BKF). SOUTH-EASTERN. Chanthaburi: Khao Soi Dao Wildlife Sanctuary, 13 July 2008, P. Phonsena, D. Chusithong & W. Chausook 6104 (BKF), 27 November 1979, T. Shimizu et al. T23830 (BKF). Trat: Taphan Hin, 3 May 1975, R. Geesink, T. Hattink & C. Phengklai 6493 (BKF, K), Koh Chang, 7 May 1974, R. Geesink, T. Hattink & C. Phengklai 6574 (BKF, K), 4 July 2001, V. Chamchumroon & C. Puff VC1166 (BKF, WU), Koh Kut, 7 April 2002, C. Phengklai 13364 (BKF), 21 October 2000, C. Phengklai 13105 (BKF). PENINSULAR. Ranong: Khlong Naka Wildlife Sanctuary, 24 December 1983, N. Fukuoka & M. Ito T35492 (BKF), Muang Ranong, Hot Spring Park, 6 December 1979, T. Shimizu et al. T26235 (BKF). Phuket: Thalang, 14 December 1979, T. Shimizu et al. T27183 (BKF). Krabi: Khao Phanom Bencha National Park, 11 July 1992, K. Larsen et al. 43306 (AAU, BKF, PSU). Phatthalung, 25 January 1958, K. Larsen & B. Hansen 649 (AAU, BKF, PSU), Tamot, 5 October 1986, P. Sirirugsa PS 1032 (BKF, PSU). Trang, Hauiyod, 3 April 2001, V. Chamchumroon VC1049 (BKF). Songkhla: Ton Sa Taw, 26 August 1995, K. Larsen et al. 46034 (AAU, BKF, PSU), Nathawi, 19 December 2006, S. Gardner ST2818 (BKF, K), 19 December 2006, S. Gardner ST2818 (BKF, K). Narathiwat: 24 March 2000, P. Puudjaa & S. Cholkulchana 717 (BKF).

Distribution: — India, Bangladesh, Myanmar, China, Japan (Ryu-kyu), Vietnam, Laos, Cambodia, Malaysia (Peninsular, Borneo), Philippines, Indonesia (Sumatra, Java, Kalimantan, Moluccas, Irian Jaya), Papua New Guinea, Thailand (Eastern, Central, South-Eastern, Peninsular).

Ecology: —Evergreen forest in shaded places at an elevation of 50‒ 900 m. Flowering in July‒October and fruiting in November‒February.

Notes: ― Lasianthus hirsutus is different from other Lasianthus by having very large and leaf-like inflorescence bracts. In the past, Lasianthus hirsutus was confused with L. cyanocarpus because they both have similar leaf-like bracts. However, Lasianthus hirsutus differs from L. cyanocarpus by having: larger leaves 13‒20 × 5‒6 cm with cuneate base (not small leaves 8–15.5 × 2.5–5 cm with cordate base); slightly unequal base (not oblique); relatively long petioles 1‒1.5 cm (not 2 mm long);conspicuous triangular stipules (not inconspicuous linear); calyx obconical densely long-hirsute (not campanulate and tomentose); drupes mosly globose, sparsely hirsute with 5 pyrenes (not drupes ovoid, sparsely pilose with 4 pyrenes);usually hirsute drupes (not tomentose).

Figures

FIGURE 3A‒O. Lasianthus werefound in Thai naturalforests:A) L. appressus;B) &C) L.attenuatus;D) L.chinensis;E) L.chrysoneurus; F) L. constrictus; G) L. cyanocarpus; H) L. ellipsoideus; I) L. ellipticus; J) L. fordii; K) L. foxworthyanus; L) & M) L. hirsutus; N) L. hispidulus and O) L. inodorus.