Lasianthus lucidus Blume (1826:997)

Napiroon, Tiwtawat, Chayamarit, Kongkanda, Dawson, Sally, Till, Walter & Balslev, Henrik, 2020, A synopsis of Lasianthus (Lasiantheae, Rubiaceae) in Thailand and two additional new species, Phytotaxa 439 (1), pp. 1-38 : 21-22

34. Lasianthus lucidus Blume (1826:997) .— Mephitidia lucida (Blume) Augustin Candolle (1830:453) . Type (lectotype designated by Zhu 2002):― INDONESIA. Java, Blume s.n. (lectotype L0000697 image!). ( Figure 3R ).

T hailand: —NORTHERN. Chiang Mai: Doi Inthanon National Park, 20 September 1995, K. Larsen et al. 46739 (AAU, BKF, PSU), 29 November 1991, R. Pooma 611 ( BKF), 10 December 1984, S. Mitsuta T48760 ( BKF), 34 October 1971, G. Murata et al. T15924 ( BKF), 29 December 1994, R. Pooma 909 ( BKF), 19 December 1965, J. F. Maxwell 96-1655 (BKF, CMUB, PSU, QBG), 14 January 1994, N. Fukuoka T62165 ( BKF). Nan: Pua, Doi Phu Kha National Park , 24 September 2000, P. Srisanga 1663 (BKF, QBG), 21 August 2001, P. Srisanga & C. Maknoi 2015 (BKF, QBG) ; Lampang: 17 December 1996, J. F. Maxwell 96-1655 (BKF, CMUB). NORTH-EASTERN. Loei: Phu Luang Wildlife Sanctuary, 19 December 2001, V. Chamchumroon VC1219 ( BKF), 13 December 1997, T. Wongprasert s.n. ( BKF), 5 December 1965, M. Tagawa, K. Iwatsuki & N. Fukuoka T1597 ( BKF) .

Distribution: —NE India, Burma, China, Vietnam, Malaysia, Philippines, Indonesia (Java, Lesser Sunda Islands, Sulawesi, Moluccas, Irian Jaya), New Guinea, Thailand (Northern, North-Eastern).

Ecology: —Evergreen forest, hill evergreen forest, and moist evergreen forest in shaded places at an elevation of 200‒1200 m. Flowering in July‒October and fruiting in November‒January.

Notes: ―In Thailand, Lasianthus lucidus is similar to L. fordii but differs by having small ovate to ovate-elliptic leaves, 4‒9 × 1.5‒3 cm (not 6‒10 × 3‒5 cm), leaf blades membranaceous (not chartaceous), calyx lobes lanceolate, 2‒3 mm long and longer than calyx tube, 1 mm long (not calyx lobes, 0.5‒0.8 mm long and mostly shorter than calyx tube, 1‒1.5 mm long). From the study of the type specimens of L. lucidus var. inconspicuous (Hook.f.) Zhu (1998:154) and L. lucidus var. flavohirsutus Zhu (2001:135) , both species are not obviously separated from L. lucidus var. lucidus . The two varieties differ slightly from the type of L. lucidus by their leaf shape (ovate-elliptic and elliptic) and a smaller number of secondary veins, which is an unstable character, as observed in the studied specimens. Lasianthus lucidus var. inconspicuous could therefore be be the same as L. lucidus var. lucidus .

Figures

FIGURE 3A‒O. Lasianthus werefound in Thai naturalforests:A) L. appressus;B) &C) L.attenuatus;D) L.chinensis;E) L.chrysoneurus; F) L. constrictus; G) L. cyanocarpus; H) L. ellipsoideus; I) L. ellipticus; J) L. fordii; K) L. foxworthyanus; L) & M) L. hirsutus; N) L. hispidulus and O) L. inodorus.