Madreallecula Kanda, 2013

Kanda, Kojun, 2013, Madreallecula Mcclevei Kanda (Coleoptera: Tenebrionidae: Alleculinae: Alleculini), New Genus and New Species of Comb-Clawed Beetle from Arizona, USA, The Coleopterists Bulletin 67 (4), pp. 587-590 : 587-588

Madreallecula Kanda , new genus ( Figs. 1–3 , 6 )

Type species: Madreallecula mcclevei Kanda , new species

Diagnosis. The combination of declivous head, moniliform antennae, and clypeus with distal angles projecting laterally separates this genus from other Alleculina . The last two characters are not found in any of the described Alleculinae in North America. Many Alleculina occurring in the Southwest United States, especially species of Hymenorus Mulsant, 1851 , superficially resemble Madreallecula in body shape and color. While the species of Hymenorus exhibit considerable variation in form and size, the antennae are always filiform ( Fig. 4 ) and the distal angles of the clypeus never project laterally. Knausia crassicornis Fall, 1931 has a similar elongate oval form but can be distinguished by its sharp posterior angles of the pronotum and slightly flabellate antennae ( Fig. 5 ). Species of Menoeceus Champion, 1888 have their head covered by their pronotum but do not possess moniliform antennae and have the clypeus more typical of other Alleculina .

Description. Body elongate oval, setose. Head declivous, partially covered dorsally by pronotum. Eyes large and prominent, notched about midway anteriorly by canthus. Vertex abruptly constricted behind eyes. Frontoclypeal suture complete. Clypeus trapezoidal, distal angles projecting laterally ( Fig. 2 ). Antennae moniliform ( Fig. 3 ), apical eight antennomeres with complex sensoria. Terminal segment of maxillary palpi large, triangular. Pronotum broad, margins complete and well-defined. Prosternum declivous, flat. Legs short. Elytra striate, epipleura terminating before

Aeanes Champion 1893 ** Narses Champion 1888 Alethia Champion 1888 * Notacula Campbell 1971 Allecula Fabricius 1801 Obesacula Campbell 1971 Amaropsis Parahymenorus

Champion 1893** Campbell 1971 Charisius Champion 1888 Phedius Champion 1888 * Diopoenus Champion 1888 Pitholaus Champion 1888 Hymenorus Mulsant 1851 * Polyidus Champion 1888 Knausia Fall 1931 * Punctacula Campbell 1971 Latacula Campbell 1971 Stenochidius

Lobopoda Solier 1835 * LeConte 1862*

Menes Champion 1888 Telesicles Champion 1888 * Menoeceus Temnes Champion 1888 Champion 1888 * Theatetes Champion 1888

apex near anterior margin of fifth visible abdominal sternite. Wings well-developed.

Etymology. The name refers to the occurrence of all known specimens only from mountains in the Madrean Sky Islands.

Figures

Fig. 1. Dorsal habitus of a paratype of Madreallecula mcclevei. Scale bar = 2 mm.

Fig. 2. Head of a paratype of Madreallecula mcclevei with clypeus emphasized. Scale bar = 0.5 mm.

Figs. 3–5. Antennae of Alleculina. 3) Madreallecula mcclevei; 4) Hymenorus sp.; 5) Knausia crassicornis. Scale bar = 1 mm.

Fig. 6. Ventral view of the aedeagus of Madreallecula mcclevei (parameres and portion of the phallobase). Scale bar = 0.25 mm.