Madrasostes simplex Paulian, 1989

Ballerio, Alberto & Maruyama, Munetoshi, 2010, The Ceratocanthinae of Ulu Gombak: high species richness at a single site, with descriptions of three new species and an annotated checklist of the Ceratocanthinae of Western Malaysia and Singapore (Coleoptera, Scarabaeoidea, Hybosoridae), ZooKeys 34 (34), pp. 77-104 : 93

Madrasostes simplex Paulian, 1989

Known distribution: East Malaysia (Sabah) ( Paulian 1979) (Fig. 1B)

Remarks. First record for West Malaysia (Selangor). Madrasostes simplex is extremely similar to M. variolosum and to M. reticulatum (Lansberge, 1887) . Madrasostes variolosum is a very common species and has a wide distribution, ranging from Thailand to Borneo (based on first author’s unpublished data) and showing a clinal morphological variation in the sculpturing of elytra, while Madrasostes reticulatum is restricted to Sulawesi and the Philippines (although the Philippine population shows intermediate characters between M. variolosum and M. reticulatum ). Madrasostes simplex can be easily distinguished from M. variolosum because of the sculpturing of elytra and pronotum and the shape of parameres.