Sibambea, ROEWER, 1917

Medrano, Miguel, Kury, Adriano Brilhante & Mendes, Amanda Cruz, 2022, Morphology-based cladistics splinters the century-old dichotomy of the pied harvestmen (Arachnida: Gonyleptoidea: Cosmetidae), Zoological Journal of the Linnean Society 195, pp. 585-672 : 639-641

SIBAMBEA ROEWER, 1917

Sibambea Roewer, 1917: 106 .

Type-species by monotypy: Sibambea rotunda Roewer, 1917 .

Discosoma Perty, 1833: 209 . Junior homonym of Discosoma Rueppell & Leuckart, 1828 (Cnidaria) .

Type-species by monotypy: Discosoma cinctum Perty, 1833 , syn. nov.

Discosomaticus Roewer, 1923: 388 . Type species automatically (ICZN Code Art. 67.8) the same as for Discosoma . Nomen novum for Discosoma Perty, 1833 , syn. nov.

Discosominana Strand, 1942: 396 . Type species automatically (ICZN Code Art. 67.8) the same as for Discosoma . Superfluous replacement name for Discosoma Perty, 1833 , syn. nov.

Etymology: Sibambea from toponym Sibambe, E c u a do r. G e n de r f e m in in e. D i s c o s o m a from Greek δίσκος (disk) + σῶμα (body). Gender neuter. Discosomaticus (gender masculine) and Discosominana (gender feminine) are derivations of the last same.

Diagnosis: DS delta-type and lenticular in lateral view ( Fig. 44C ); differs from other genera of the subfamily by the combination of the following features: (1) armature of areas of mesotergum absent or extremely reduced ( Fig. 46D ); (2) coxa IV with basal thickening ( Fig. 46C ); penis with rectangular and short VP with concave apical margins ( Fig. 43C ); and (3) stylus with remarkable stylar barbs perpendicular to a short wattle ( Fig. 43D ).

Included species: Sibambea rotunda Roewer, 1917 , S. cincta ( Perty, 1833) comb. nov. and S. waorani Medrano, Kury & Mendes sp. nov.

KEY TO THE SPECIES OF SIBAMBEA

1. DS with marginal yellow ring ramification to the mesotergum but not occupying considerably the areas of abdomen ( Fig. 44A, B ). Coxa IV with coloured tubercles, a proximal row well defined with five or six tubercles boarding the DS ( Fig. 44C ) ............................................................................................... S. rotunda

1’. DS with marginal yellow ring with entire inner borders (not branched into mesotergum, Fig. 42A ) or a large spot occupying almost the entire DS ( Fig. 46A ). Coxa IV smooth without any coloured spot ( Figs 42C , 46D )......................................................................................................................................................2

2. Light coloration of DS with a hollow area mainly in the mesotergum, free tergites coloured shaping the ring of DS ( Fig. 42A, C, D ). Fixed finger of chelicera with a row of well marked teeth ( Fig. 43E ). Dorsal process of the stylus apically subdivided in five lobes ( Fig. 43D ) ....................................................... S. cincta

2’. DS with a large yellow spot occupying almost the entire DS ( Fig. 46A–D ). Fixed finger of chelicera with a basal tooth ( Fig. 47E ). Dorsal process of the stylus apically entire ( Fig. 47A, B ) ................. S. waorani

Figures

Figure 42. Sibambea cincta, male (MPUJ_ENT 644772). A, dorsal view. B, panoramic dorsal view. C, lateral view. D, posterior view. E, ventral view. Scale bars: 1 mm.

Figure 43. SEM images of some structures of Sibambea cincta, male (ICN-Ao- 972). A–D, penis: A, dorsal view; B, lateral view; C, ventral view; D, detail of stylus and dorsal process of glans. E, left cheliceral hand, dorsal view. F, left basichelicerite, dorsal view. G, left pedipalpus, ectal view. H, same, mesal view. I, same, detail of tarsus, ventral view. Scale bars: 50 μm (A–C), 10 μm (D), 200 μm (E, G, H), 100 μm (F, I).

Figure 44. Sibambea rotunda, male (CAS AK 003). A, dorsal view. B, lateral view. C, panoramic dorsal view. Scale bars: 1 mm.

Figure 46. Sibambea waorani, male holotype (QCAZ AK 156). A, dorsal view. B, panoramic dorsal view. C, posterior view. D, lateral view. Scale bars: 1 mm.

Figure 47. SEM images of some structures of Sibambea waorani, male. A–D, penis (FMNH AK 121g): A, dorsal view; B, lateral view; C, ventral view; D, detail of stylus and dorsal process of glans. E–H, male (FMNH AK 124a): E, left cheliceral hand, dorsal view; F, left basichelicerite, dorsal view; G, left pedipalpus, ventral view; H, same, mesal view. Scale bars: 50 μm (A–C), 20 μm (D), 200 μm (E, G, H), 100 μm (F).