Norrbomia paragravis, Papp, László, 2017

Papp, László, 2017, Twenty New Species Of Fifteen Genera Of The Afrotropical Sphaeroceridae (Diptera), Acta Zoologica Academiae Scientiarum Hungaricae 63 (4), pp. 377-427 : 380-383

Norrbomia paragravis sp. n.

( Figs 8 –24)

Holotype: male ( BMSA), sweepingcattle/horsedung – RSA: FreeState , Harrismith, Geluhsdalfarmat: 27° 54’ 7.05’’S, 29° 23’ 31.9’’E, 9–10.xi.2009, A. H. Kirk- Spriggs – [blue] EntomologyDept. NationalMuseum, P.O.Box 266, Bloemfontein 9300, SouthAfrica – BMSA (D) 13395.

Paratypes: 8 males 10 females ( BMSA): datasameasforholotype ; 1 male ( HNHM): EasternCapeProv., ShamwariGameReserve, onelephantdung, Jan 11, GPS10, S33° 24’ 47.0’’ E26° 05’ 45.0’’, 301 m, No. 14, leg. L. Papp & M. Földvári.

Size (inmm): bodylength 2.13 (holotype), 1.65–2.75 (paratypes); winglength 2.17 (holotype), 1.67–2.46 (paratypes); wingwidth 0.80 (holotype), 0.71–0.92 (paratypes).

Description. Male. Frontaltrianglewithsomeshine (lessdusted), 4 pairsofcomparativelyshortinterfrontalsetae, shorterthan 0.10 mm. Postocellars ca. halfocellars length. Genawithsubtriangularbareareabeloweye, verynarrownearvibrissa. Arista rathershort: 0.35–0.45 mm (0.40 mmonholotype). Mesonotumthicklymicrotomentose. 1+3 dorsocentralpairs, acrostichalsetaesparseintworegularrows. Katepisternumwith alargeventralcaudalshinyspotreachingonlymidofkatepisternum. Wingmembrane lightbrownish, veinslightbrown. Legsdark, microtomentose, exceptforekneeandmid basaltarsomeres, darkyellowish. Midtibiawithsetaeasfollow: 2 strongventro-apical, 2 Figs 19–24. Norrbomia paragravis sp. n., female postabdomen and genitalia. 19 = preabdomi- nal sternites, ventral view, 20 = sternite 8, ventral view, 21 = tergite 8 and sternite 8, lateral view (arrow: positioning to body axis), 22 = epiproct and cerci, dorsal view, 23 = hypoproct, ventral view, 24 = spermathecae. Scale bar: 0.4 mm for Fig. 19, 0.2 mm for Figs 20–23, 0.1

mm for Fig. 24 anteriorapical, amedium-longanterodorsalat 2/3, andapreapicalanterodorsalaswellas a posterodorsal at 5/6. Hind tibia with a large (0.21 mm) long curved ventral spur, with a long (0.20–0.22 mm) anterodorsalsetaat 7/9 anda 0.20–0.21 mmlongdorsalpreapicalseta.

Preabdominaltergitesnotparticularlysmall, withthicklightmicrotomentum: sternite 4 c. 1/4 as broad as opposite tergite, sternite 3 c. 1/5 as broad as opposite tergite. Sternite 5 ( Fig. 8) shield-shaped, aboutaslongasbroad (28: 30) withacaudalsublateralpairof processes 0.06 mm long, out of the plane of sternite though not perpendicular to it. Sides of sternite 6 and 7 ratherlarge. Postabdominalscleritesmoderatelyshiny. Hypandrium ( Fig. 9) rather long, medial part (rod, continued to apex of caudal process) 0.37 mm, arms (from rodtolateralapex) 0.21 mm. Subepandrialsclerite ( Fig. 14) withatrapezoidbaseandwith a pair of digitiform ventral processes. Cercus ( Figs 10, 15–16) bilobed, medial lobe with abluntcranialandcaudalprocesseach, laterallobesubtriangular. Surstylus ( Figs 11–12) actually short, very broad, with rounded apex, with rather few setae, but two apical teeth. Phallapodeme ( Fig. 13) ratherhighforitsdiameter. Basiphallus ( Fig. 13) withanextremely large (butnotthick) anteriorlycurvedcaudalprocess. Distiphallus ( Fig. 18) withcomplex shape: adorsallycurvedblacksub-basalprocessandananteriorlycurvedbluntmedial process; apexnotsharp. Postgonitebroad ( Fig. 17) medialpartbroadlylengthened, actually notventralprocess, sinceventraledgebroaderthansub-basalpartbutapexnotprotruding.

Female. Sternite 4 (Fig. 19) withpeculiarshape, mostlymembranous, apairofelongatedlateralsclerites, threepairsofroundmesialscleritessclerotised. Sclerites 6 to 8 strongly shiny. Sternite 8 (Figs 20–21) divided in two parts, deeply protruding (in some specimensalmostperpendicularly); ventralpartwithanarrowapicallyroundedprocess. Hypoproct (Fig. 23) shield-shaped, with 2 pairsofverylongsetae. Epiproct (Fig. 22) small, withapairofbasal, anteriorlydirectedprocesses, bodyofepiprocttriangular, with 3 pairs ofpartiallyasymmetricalsetae. Cercus (Fig. 22) regular, withtwopairsoflongapicalsetae andseveralpairsofmediumtolongsetaesubapicallyandlaterally. Spermathecae (Fig. 24) notglobular, definitelyshorterthanbroadwithacylindricalmesalpartandalateralcavity; nohairsorscalesonspermathecae.

Comments. N. paragravis sp. n. isaquitepeculiarspecies, althoughithas all diagnostic features of Norrbomia listed by PAPP (1988: 394). In PAPP’s (1988) key, itmeetsaconflictatcouplet 1, mesonotumthicklymicrotomentose. Our specieshas 4 (1+3) dorsocentralpairs. Thehighnumberofstrongdorsocentral setae is shared with N. gravis , but it differs from N. gravis (Adams) due to its mat (microtomentose) mesonotum. N. gravis hasmostly 5 pairsofdorsocentrals. Asforthemalegenitalia, thenewspeciesiseasilyseparatedfromthe remainingspeciesduetoitsbilobedcercus.

Etymology. Thespecificepithetofthisnewspecies (‘paragravis’ = besidegravis) referstoitsrelationto N. gravis (Adams) .