Anyphaena triangularis, Rivera-Quiroz & Álvarez-Padilla, 2023

Rivera-Quiroz, F. Andrés & Álvarez-Padilla, Fernando, 2023, Integration or minimalism: twenty-one new species of ghost spiders (Anyphaenidae: Anyphaena) from Mexico, European Journal of Taxonomy 865, pp. 1-94 : 20-24

publication ID	https://doi.org/ 10.5852/ejt.2023.865.2097
publication LSID	lsid:zoobank.org:pub:845BAE1A-0E6E-484B-BE6D-17F866777B51
DOI	https://doi.org/10.5281/zenodo.7887410
persistent identifier	https://treatment.plazi.org/id/03C8E927-FFA7-FF87-FDD9-FEF0FE9C6225
treatment provided by	Felipe
scientific name	Anyphaena triangularis
status	sp. nov.

Treatment

Anyphaena triangularis sp. nov.

urn:lsid:zoobank.org:act:99F21C7F-3CC1-413F-9269-57931511CE81

Figs 10–12 View Fig View Fig View Fig , 52 View Fig

Differential diagnosis

The epigynal plate in A. triangularis sp. nov. is broad, more than two thirds of the epigastric furrow length, anteriorly projected and ventrally distended, epigynum anterior projection with two deep comma-shaped pits ( Figs 10C, F View Fig , 11E–F View Fig , 12G View Fig (arrow), H). Similar epigyna are found in A. urieli sp. nov., but they differ by the flatter anterior projection and copulatory duct paths. Males of A. triangularis sp. nov. are differentiated from those of A. urieli sp. nov. by their larger cymbium basal projection ( Figs 11C View Fig , 12C View Fig (arrow)), ventral tegular projection retrolateral margin without a translucent section, median apophysis distal hook smaller, the RTA posterior branch wider, and the coxa II tubercle larger ( Figs 11A–D View Fig , 12A– F View Fig ).

Etymology

The species epithet refers to the triangular shape of the epigynal plate in ventral view.

Material examined

Holotype MEXICO • ♀; San Luis Potosi, Xilitla City, Las Pozas ; 21.39722° N, 98.99388° W; alt. 662 m; 10–15 Jun. 2012; Aracnolab team leg.; tropical wet forest fragment; BEAT; CNAN-T01541 . GoogleMaps

Allotype MEXICO • ♂; same collection data as for holotype; CNAN-T01520 . GoogleMaps

Paratypes MEXICO • 1 ♀; same collection data as for holotype; LUP; CNAN-T01579 GoogleMaps • 1 ♀; same collection data as for holotype; CNAN-T01580 GoogleMaps • 1 ♂; same collection data as for holotype; CNAN-T01577 GoogleMaps • 1 ♂; same collection data as for holotype; CNAN-T01578 . GoogleMaps

Additional material

MEXICO • 3 ♂♂; San Luis Potosi, Xilitla City, Las Pozas ; 21.39722° N, 98.99388° W; alt. 662 m; 10–15 Jun. 2012; Aracnolab team leg.; tropical wet forest fragment; BEAT GoogleMaps • 1 ♀; same collection data as for preceding; ANYM085 GoogleMaps • 1 ♂; same collection data as for preceding; CRP GoogleMaps • 1 ♀; same collection data as for preceding; ANYM086 GoogleMaps • 1 ♀; same collection data as for preceding; ANYM087 GoogleMaps • 3 ♂♂; same collection data as for preceding; LUD GoogleMaps • 2 ♀♀; same collection data as for preceding; ANYM090 GoogleMaps • 2 ♂♂; same collection data as for preceding; ANYM091 GoogleMaps • 4 ♂♂; same collection data as for preceding; ANYM092 GoogleMaps • 5 ♂♂; same collection data as for preceding; LUP GoogleMaps • 1 ♀; same collection data as for preceding; ANYM088 GoogleMaps • 1 ♀; same collection data as for preceding; ANYM089 GoogleMaps • 1 ♀; same collection data as for preceding; 27–31 Jul. 2011; CRP; AR_002; GenBank: ON619642 . GoogleMaps

Description

Female

Total length 6.9. Carapace yellow, ocular area slightly darker, pattern with two darker longitudinal brown parallel bands around cephalic area and fovea, clypeus dark yellow ( Fig. 10A, D View Fig ). Sternum surface white, intercoxal triangles present on all legs. Labium yellow, white at tip, longer than wide. Endites yellow, rectangular, slightly broader at tip ( Fig. 10C View Fig ). Chelicerae dark yellow, paturon dorsum without pattern ( Fig. 10B, D View Fig ), promargin with five teeth, retromargin with eleven to twelve denticles. Leg coloration: yellow covered with darker patches from femora to tarsi. Abdomen yellow, hirsute, dorsal surface delineated with light brown pattern and several chevrons over posterior half, anterior central half with darker median band, lateral surfaces light brown pattern only at top, lateral sides and ventral surface white, tracheal spiracle at middle ( Fig. 10A–C View Fig ). Copulatory openings under semicircular and sclerotized hood. Hood with straight edge slightly projected at center. Atrium short, lateral borders sinuous, extended in posterior region ( Figs 11E–F View Fig , 12G–H View Fig ). Copulatory ducts sclerotized, short, entering spermathecae on anterior surfaces. Seminal receptacles rounded and located in anterior third of copulatory ducts. Fertilization ducts short, straight, and emerging from lateral surfaces of spermathecae ( Figs 11E–F View Fig , 12G–F View Fig ). Cephalothorax length 2.93, thoracic width 2.1, cephalic width 0.97. Clypeus height 0.09. Eye diameters: AME 0.07, ALE 0.13, PME 0.12, PLE 0.12. Eye interdistances: AME– AME 0.07, AME–ALE 0.04, ALE–PLE 0.06, PME–PME 0.12, PME–PLE 0.09. Femur lengths: I 3.38, II 3.05, III 2.29, IV 3.43. Leg spination: femur I d1-1-1, p0-2-1, r0-1-1. Tibia I v2-2-0, p1-1-1, r1-1-1. Metatarsus I v2-2-0, p1-1-1, r1-1-1. Femur II d1-1-1, p0-1-1, r0-1-1. Tibia II v2-2-0, p1-1-1, r1-1-1. Metatarsus II v2-2-0, p1-1-1, r1-1-1. Femur III d1-1-1, p0-1-1, r0-1-1. Tibia III v2-2-2, p0-1-1, r0-1-1. Metatarsus III v2-2-2, p1-1-2, r1-1-2. Femur IV d1-1-1, p0-0-1, r0-0-1. Tibia IV v2-2-2, p0-1-1, r0-1-1. Metatarsus IV v2-2-2, p1-1-2, r1-1-2.

Male

Total length 6.9. Cephalothorax and abdomen coloration as in female ( Fig. 10E View Fig ). Retromargin of chelicerae with seven to eight denticles. Coxa III tubercle roughly rectangular with posterior apical spine. Coxa IV with a long, curved spur, located at retrolateral basal corner ( Fig. 12E–F View Fig ). Ventral tegular apophysis bifurcated ( Fig. 12A View Fig (arrow)). Median apophysis hook-shaped in ventral view. Embolus long, translucent and filiform ( Figs 11A, D View Fig , 12A, D View Fig ). RTA anterior branch curved with sclerotized margin, posterior branch of RTA large, distal edge with translucent keel ( Figs 11A, C–D View Fig , 12A, C–D View Fig ). Prolateral apophysis of palpal tibia small, conical ( Figs 11B–C View Fig , 12B–C View Fig ). Pedipalp tibia proportions longer than wide, stridulatory ridges extending from ventral branch of median tegular apophysis to tibial distal margin ( Figs 11A–B, D View Fig , 12A–B, D View Fig ). Cephalothorax length 2.78, thoracic width 2.15, cephalic width 0.97. Clypeus height 0.11. Eye diameters:AME 0.09, ALE 0.16, PME 0.16, PLE 0.15. Eye interdistances: AME–AME 0.05, AME–ALE 0.04, ALE–PLE 0.05, PME–PME 0.1, PME–PLE 0.09. Femur lengths: I 3.52, II 3.19, III 2.38, IV 3.57. Leg spination as in female except: femur IV r0-1-1.

Variation

Females (N =10): total length 6.94 (± 0.36), cephalothorax length 2.73 (± 0.11), thoracic width 2.06 (± 0.07), cephalic width 1.12 (± 0.06), femur I 3.32 (± 0.12). Males (N=10): total length 6.66 (± 0.26), cephalothorax length 2.74 (±0.1), thoracic width 2.19 (± 0.1), cephalic width 1.06 (± 0.06), femur I 3.6 (± 0.06).

Distribution

This species is found in primary tropical wet forest fragments ( Fig. 52 View Fig ).

Natural history

Most specimens were collected over vegetation by beating, direct searching and cryptic search on the ground.