Anopinella phillipsae Brown and Adamski

Brown, John W. & Adamski, David, 2003, Systematic revision of Anopinella Powell (Lepidoptera: Tortricidae: Euliini) and phylogenetic analysis of the Apolychrosis group of genera, Zootaxa 200, pp. 1-94 : 35-36

Anopinella phillipsae Brown and Adamski , new species

Figs. 25 , 48 , 78

Diagnosis. Anopinella phillipsae is most similar to A. larana in facies and genitalia. Both have bronzy, recurved scales that diminish the definition of the triangular costal blotch, although our specimens of A. larana are rubbed so this feature is not particularly conspicuous. Both species have the large, bent, unmodified uncus; a gnathos typical of the Fana Species Group; extremely short socii; and comparatively long, narrow, apically attenuate valvae with a costa that is slightly upturned at its apex. The elongate­oval sclerotized mesal patch at the lamella postvaginalis in the female genitalia is also similar between A. phillipsae and A. larana . The conspicuous patches of shiny recurved scales in the basal ca. 0.65 of the forewing of A. phillipsae may represent an autapomorphy for this species, although these may be present in A. larana as well.

Description. Head: Frontoclypeus and vertex pale brown; labial palpus with outer surface of segments I­II brown intermixed with few dark brown and pale brown scales, segment III pale brown, inner surface pale brown. Antenna with scape pale brown; basal 8­10 flagellomeres reddish brown; distal flagellomeres gray.

Thorax: Tegula brown basally, pale brown distally, mesonotum pale brown. Forewing (Fig. 78) length 6.9­8.0 mm (n = 8); incomplete basal fascia and costal blotch olive­green intermixed with reddish brown and brown scales, merging posteriorly near cubitus; basal fascia and costal blotch separated anteriorly by an incomplete oblique pale band of pale brown intermixed with pale olive­green and pale reddish brown; basal ca. 0.65 with most scales slightly upturned apically, reflecting bright yellow­bronze, with several reddish brown and gray patches; costal blotch with a small subrectangular white spot near posterior end; area of wing between DC and submargin white or pale brown encircling a suffused brown ocellus; apical area brown intermixed with white and reddish brown; submarginal area dark brown intermixed with dark reddish brown. Fringe alternating reddish brown and gray. Hindwing pale grayish brown.

Abdomen: Male genitalia ( Fig. 25 ; drawn from DA slide 4563; n = 2) with uncus bent at basal 0.25. Socius short, digitate, ca. 0.14 length of gnathos arms. Gnathos arms slightly widened basally, distally enlarged forming two elongate, decumbent lobes with entire margins; lobes connected dorsally by a hood­shaped flange. Valva sparsely setose from cucullus to basal ridge; costa nearly straight in basal two­thirds, slightly angled dorsad in distal one­third, ending in a pointed apex; sacculus nearly straight; postsacculus nearly straight, with a broadly rounded apicoventral angle. Phallus simple, long; vesica densely microtrichiate. Female genitalia ( Fig. 48 ; drawn from JWB slide 1196; n = 6) with elongate sclerotized mesal patch at lamella postvaginalis; eighth tergum with a deeply invaginated crescent­shaped flange ( Fig. 48 insert); ostium narrow, shallowly U­shaped, ca. 0.33 width of seventh sternum at widest point. Antrum slightly elongate, parallel­sided; ductus bursae long, slender, gradually widening anteriorly, with sparse spicules on anterior 0.33 of side bearing accessory bursae; inception of accessory bursae ca. 0.75 distance from ostium to corpus bursae. Corpus bursae subspherical, sparsely spiculate on ca. 0.33 of side opposite ductus seminalis.

Holotype, ɗ, Costa Rica , Cartago Province, Paraíso, P.N. Tapantí­Macizo de la Muerte, 300 m SE del Río Porras, 1660 m, 1 May 2000, R. Delgado. Deposited in INBio.

Paratypes (1ɗ, 6Ψ). COSTA RICA : Cartago Province: Paraíso, P.N. Tapantí­Macizo de la Muerte, 300 m SE del Río Porras, 1660 m, Jan 2000 (2Ψ), R. Delgado (INBio, USNM), Nov 2001 (1Ψ), R. Delgado (INBio). Paraíso, P.N. Tapantí­Macizo de la Muerte, R. Grande de Orosi desde Admin. hasta Sendero la Pava, Nov 1995, (1Ψ), G. Mora (INBio). Paraíso, P.N. Tapantí­Macizo de la Muerte, 300 m N del Mirador, 1830 m, Jul 2000 (1ɗ), R. Delgado (USNM). Ref. Nac. Fauna Silv. Tapantí, 1250 m, Aug 1991 (1Ψ), G. Mora (INBio). Tapantí, 1200­1700 m, 20 Aug­15 Sep 1999 (1Ψ), V. Becker (VBC).

Etymology. The species name honors Eugenie Phillips, our colleague in Costa Rica who has facilitated visits to that country and the study of material deposited in Instituto Nacional de Biodiversidad.

Figures

FIGURES 25 ­ 26. Male genitalia of Anopinella, valvae spread, aedeagus removed. 25, A. phillipsae; 26, A. larana.

FIGURES 48 ­ 49. Female genitalia of Anopinella, posterior end up. 48, A. phillipsae, with insert of tergum eight; 49, A. larana.