Anopinella triquetra (Walsingham, 1914)

Brown, John W. & Adamski, David, 2003, Systematic revision of Anopinella Powell (Lepidoptera: Tortricidae: Euliini) and phylogenetic analysis of the Apolychrosis group of genera, Zootaxa 200, pp. 1-94 : 11-12

Anopinella triquetra (Walsingham, 1914)

Figs. 2 , 35 , 57

Tortrix triquetra Walsingham 1914: 291 .

Seticosta triquetra ; Razowski 1986: 22.

Anopinella triquetra ; Powell et al. 1995: 142.

Diagnosis. As discussed above, A. triquetra is most similar to A. isodelta . The two can be distinguished by the characters described in the diagnosis of A. isodelta . Putative autapomorphies for A. triquetra include the following: male genitalia with the posteroventral part of the gnathos lobes greatly expanded, and eighth tergum of the female with a weak lateral sinus at middorsum flanked by two oblique flanges, resulting an a shallowly U­shaped ridge ( Fig. 35 insert).

Redescription. Head: Frontoclypeus white intermixed with few pale gray scales; vertex pale gray intermixed with reddish brown; labial palpus with outer surface brown intermixed with reddish brown and few white scales, inner surface white. Antenna with scape brown intermixed with some reddish brown and pale brown scales; basal 6­10 flagellomeres reddish brown; distal flagellomeres gray.

Thorax: Tegula and mesonotum pale brownish yellow intermixed with dark brown and reddish brown. Forewing (Fig. 57) length 7.0­ 9.5 mm (n = 10); incomplete basal fascia and costal blotch brown intermixed with pale brown and few reddish brown and dark brown scales, separated by a pale, oblique band of pale brown intermixed with pale reddish brown scales; some specimens with more white scales demarcating costal blotch, recurved from costa through subapical area to tornus, encircling an elliptical ocellus, except on posterior part; elliptical ocellus usually reduced, absent in some specimens; costal blotch subtriangular, with a small, subrectangular, white spot near posterior end; apical area white intermixed with brown; submarginal area with gray patches separated by thin, reddish brown bars. Fringe with outer half brown and/or gray, inner half reddish brown and brownish gray. Hindwing pale brown, gradually darkening to apex, or brown.

Abdomen: Male genitalia ( Fig. 2 ; drawn from USNM slide 68841; n = 1) with uncus curved from base. Socius relatively wide, digitate, ca. 0.65 length of gnathos arms. Gnathos greatly expanded posteroventrally, forming two free, laterally flattened lobes, each with outer margin sparsely serrate. Valva densely setose from cucullus to basal ridge; costa nearly straight from subbasal curve to apex; ventral edge with basal 0.66 weakly, evenly curved, then recurved to apicoventral angle; cucullus elongate, acuminate, gradually widening to mid­length, recurved to lateroventral margin, forming a protuberant apex. Phallus simple; vesica densely microtrichiate. Female genitalia ( Fig. 35 ; drawn from USNM slide 68858; n = 4) with weakly sclerotized, elliptical mesal patch at lamella postvaginalis; eighth tergum with a short lateral median sinus flanked by two small, oblique flanges ( Fig. 35 insert); ostium broadly U­shaped, about 0.33 width of seventh sternum, ventral rim narrowly sclerotized. Ductus bursae long, slender; inception of accessory bursa ca. 0.5 distance from ostium to corpus bursae. Corpus bursae large, subspherical, spiculate on 0.25 side bearing ductus seminalis.

Holotype, Ψ, Guatemala, Baja Vera Paz, San Geronimo, 1879­80, G. C. Champion [head missing]. Deposited in BMNH.

Additional Specimens Examined. GUATEMALA: Volcan Santa Maria, Jun­Jul [no year] (1ɗ, 20Ψ), Schaus and Barnes Collection (USNM).

Remarks. We examined two additional females from Honduras (Morazán, Tegucigalpa, La Tigre, 1800 m, 24 Aug 2000, V. Becker, VBC) in which the genitalia are similar to those of A. triquetra , with an inverted U­shaped flange on the eighth tergum. However, the forewing pattern is conspicuously darker in these specimens, without the broad, pale, subterminal area. Hence it is possible that these specimens represent an undescribed species near A. triquetra . We also examined two females from Costa Rica ( Cartago Province, Paraíso, P.N. Tapantí­Macizo del la Muerte, Est. Quebrada Segundo, 1300 m, Mar 2001, R. Delgado, INBio; and Puntarenas Province, La Amistad, Sector Altamira, Cerro Biolley, A.C. Amistad, 1800 m Jan 1994, R. Delgado, INBio) that are similar to A. triquetra but differ slightly in the genitalia from those of the series from Guatemala. Because we are uncertain if they are conspecific with A. triquetra we treat them as A. nr. triquetra .

Figures

FIGURES 1 ­ 2. Male genitalia of Anopinella, valvae spread, aedeagus removed. 1, A. isodelta; 2, A. triquetra.

FIGURES 34 ­ 35. Female genitalia of Anopinella, posterior end up. 34, A. isodelta; 35, A. triquetra, with insert of tergum eight.