Hlavaciellus cirrus, Jałoszyński, Paweł, 2011

Jałoszyński, Paweł, 2011, Hlavaciellus Jałoszyński, 2006: eleven new species, detailed morphology and systematic position within Cephenniini (Coleoptera, Staphylinidae, Scydmaeninae), Zootaxa 2763, pp. 1-33 : 5-7

Hlavaciellus cirrus sp. n.

( Figs. 2 , 5 , 15–17 )

Type material. Holotype: MALAYSIA: ɗ, "E. MALAYSIA: Sarawak \ Gn. Matang, 20 km E \ Kuching, 850 m, 25.V.1994 \ submontane for. #10a \ Löbl & Burckhardt" [white, printed], " HLAVACIELLUS \ cirrus m. \ det. P. JAŁOSZYŃSKI, '09 \ HOLOTYPUS " [red, printed] ( MHNG).

Diagnosis. Head in males medially with two large but indistinctly delimited and flat tubercles accompanied posteriorly by median tuft of long, curly setae.

Description. BL 1.98. Body of male ( Fig. 2 ) moderately convex, pigmentation reddish-brown, vestiture yellowish.

Head ( Figs. 5 , 17 ) large, HL 0.28, HW 0.50; vertex long and regularly convex, delimited from frons by pair of very shallow and diffused lateral impressions running anterad and towards middle from posterior margin of each eye; frons slightly elevated in relation to anterior part of vertex, with median tuft of long and curly setae and median pair of indistinct, diffused and flattened tubercles; supraantennal tubercles prominent, distinctly separated from frons and vertex; eyes large, strongly convex and coarsely faceted. Punctures and setae as in Fig. 5 . Antennae moderately long and slender, reaching 0.66 of BL, gradually thickened up to antennomere X, uniformly covered with thin, long, dense and suberect setae, AnL 1.30, antennomere I 1.4x as long as broad; II distinctly narrower and much shorter than I, only 1.1x as long as broad; III slightly broader and distinctly longer than II, 1.2x as long as broad IV slightly longer and broader than III, 1.3x as long as broad; V barely noticeably broader but not longer than IV, nearly 1.3x as long as broad; VI distinctly longer and broader than V, 1.3x as long as broad; VII–VIII nearly equal in length and width, each slightly larger than VI, nearly 1.2x as long as broad; IX slightly larger than VIII, nearly as long as broad; X slightly larger than IX, about as long as broad; XI distinctly narrower than X, slightly shorter than IX–X together.

Pronotum in dorsal view subrectangular with broadly rounded anterior margin, nearly equally broad from base up to anterior fourth, PL 0.60, PW 0.88; sides strongly rounded in anterior fourth, then feebly (barely noticeably) concave; hind angles slightly sharp but not acute; posterior margin shallowly bi-emarginate; ante-basal pits very shallow and diffused, each located much closer to posterior than to lateral margin of pronotum; lateral edges indistinctly swollen. Pronotal disc glossy, in middle covered with very fine and shallow punctures separated by spaces 1.5– 2 x as wide as puncture diameters, punctures close to margins of pronotum are larger and denser, those covering subtriangular area between each hind angle and ante-basal pit are slightly coarse. Setae thin, long, moderately dense, suberect to erect.

Elytra oval, relatively long, moderately convex and gradually lowering towards apices in posterior half, broadest near anterior third, EL 1.10, EW 0.88, EI 1.26; basal pit on each elytron small but distinct, located very close to scutellum; subhumeral lines very distinct, as long as 0.36x EL, only slightly divergent, each developed as sharp border between higher humeral region and lower adsutural area; apices of elytra separately rounded. Punctures on basal third of each elytron small but more distinct and denser than those on median part of pronotal disc, gradually reducing in diameter and depth toward lateral margins and apices; setae similar to those on pronotum but slightly thicker and more erect. Hind wings not studied.

Legs slender and long, all tibiae straight.

Metaventrite with moderately deep postmesocoxal impressions, occupying about 2/3 of its length.

Aedeagus ( Figs. 15, 16 ) very slender, AeL 0.50; median lobe with triangular and pointed apical part; internal armature nearly symmetrical; parameres in lateral view extremely broad, with apices curved more towards apex of median lobe than dorsally.

Female. Unknown.

Distribution. East Malaysia: Borneo, Sarawak.

Etymology. The Latin noun cirrus , meaning a lock, curl, ringlet, or tuft of hair, was chosen to underline the most remarkable diagnostic character of the new species, the curly tuft of setae on the frons.

Remarks. Although the aedeagus of H. cirrus is unremarkable and very similar to copulatory organs of several other species described in this paper, the modification of the head is unique and remotely similar only to that of H. vampirus . However, the latter species has only a pair of large tubercles on the frons, without the remarkable median tuft of curly setae, and the apex of the aedeagus in that species in lateral view is much more slender and strongly recurved (stout and nearly straight in H. cirrus ).

Figures

FIGURES 1 – 3. Dorsal view of holotypes of Hlavaciellus, males. 1, H. peninsularis sp. n. 2, H. cirrus sp. n. 3, H. angustifrons sp. n. Scale bars: 1 mm.

FIGURES 4 – 9. Modifications of head in frontal view. 4, Hlavaciellus peninsularis sp. n. 5, H. cirrus sp. n. 6, H. angustifrons sp. n. 7, H. decoratus sp. n. 8, H. smetanai sp. n. 9, H. bifoveatus sp. n. Scale bars: 0.1 mm.

FIGURES 13 – 23. Aedeagus in ventral (13, 15, 18, 20, 22) and lateral (14, 16, 19, 21, 23) views, and head of male in lateral view (17). 13, 14, Hlavaciellus peninsularis sp. n. 15 – 17, H. cirrus sp. n. 18, 19, H. angustifrons sp. n. 20, 21, H. decoratus sp. n. 22, 23, H. smetanai sp. n. Scale bars: 0.2 mm (13 – 16, 18 – 23), 0.1 mm (17).