Eumenes verticalis Say, 1824
publication ID |
https://doi.org/ 10.11646/zootaxa.4459.1.1 |
publication LSID |
lsid:zoobank.org:pub:633A9498-FCC6-4439-BE33-A45B9E51B550 |
DOI |
https://doi.org/10.5281/zenodo.5986390 |
persistent identifier |
https://treatment.plazi.org/id/03C5878F-FFEE-F31E-FF30-9339D7F2F8BA |
treatment provided by |
Plazi |
scientific name |
Eumenes verticalis Say, 1824 |
status |
|
Eumenes verticalis Say, 1824 View in CoL
Eumenes verticalis Say, 1824 , 2: 346.
Type data: Type destroyed.
Type locality: “ Pennsylvania ”
de Saussure, 1852, 1: 41.
Cresson, 1887: 287 (cat.).
Dalla Torre, 1894, 9: 33 (cat.).
Dalla Torre, 1904, 19: 25 (cat.).
Bequaert, 1928, 101: 1004.
Bequaert, 1938, 33: 61 (key), 64, 70 (type lost).
Bequaert & Taylor, 1942, 42: 68 (list).
Bequaert, 1944, 71: 78 (key), 86.
MacLachlan, 1980, 53: 618, 620 (key).
Buck et al. 2008, 5: 44 (key), 155.
Eumenes coloradensis Cresson, 1875 , 5: 717. REVISED STATUS.
Type data: Lectotype female at ANSP (examined).
Type locality: Colorado ( USA).
Dalla Torre, 1894, 9: 21 (cat.).
Dalla Torre, 1904, 19: 21 (cat.).
Tucker, 1909, 22: 286.
Cresson, 1916, 1: 104 (designation of Lectotype).
Isely, 1917, 10: 348 (key), 359.
Bequaert, 1938, 33: 65, 69 (note on types).
Bequaert, 1944, 71: 78 (key), 81.
Eumenes (Alpha) cruciferorum Viereck, 1908 , 33: 388. REVISED STATUS.
Type data: Lectotype female (by present designation) and Paralectotype male at KUNHM (examined).
Type locality: “ Thomas’ Ranch, Oak Creek Canon , 6000 ft., 20 miles southwest of Flagstaff , Coconino Co., Arizona ”
(USA).
Eumenes cruciferorum ; Bequaert, 1938, 33: 69 (note on types).
Bohart, 1951: 885.
Eumenes (Alpha) enigmatus Viereck, 1908 , 33: 389. NEW STATUS.
Type data: Lectotype male (by present designation) and Paralectotype male at KUNHM (examined).
Type locality: “Thomas’ Ranch, Oak Creek Canon, 6000 ft., 20 miles southwest of Flagstaff, Coconino Co., Arizona”. Eumenes enigmatus ; Bequaert, 1938, 33: 69 ( enigmaticus [!]; note on type).
Bohart, 1951: 885.
Eumenes stenogaster Isely, 1917 View in CoL , 10: 353. NEW STATUS.
Type data: Holotype female at USNM (n° 21379) (examined).
Type locality: “ White Mts , Novo Mexico ; Rio Ruidoso ; about 6500 ft.” ( USA) .
Bequaert, 1938, 33: 70 (note on types).
Bohart, 1951: 885.
Eumenes tricinctus Isely, 1917 , 10: 361. REVISED STATUS.
Type data: Holotype female at USNM (n° 21383) (examined).
Type locality: “ Oregon ”.
Bequaert, 1938, 33: 70 (note on type).
Eumenes verticalis View in CoL var. (or subsp.) neoboreus Bequaert, 1944 View in CoL , 71: 76, 87, NEW STATUS
Type data: Holotype male at MCZ (n° 33185) (examined). Three paratypes male at ROM (examined).
Type locality: “ Canada... Quebec: La Trappe ” (Holotype) ; “ Canada: Ontario (paratypes) .
Eumenes verticalis tricinctus ; (4) 24 (9): 315.
Bohart, 1951: 885 (cat.)
Krombein, 1979: 1508 (cat.).
MacLachlan, 1980, 53: 618.
Ruiz C. et al., 1993, 88: 87 (list).
Rodríguez-Palafox, 1996: 479 (list).
Eumenes verticalis verticalis ; Bohart, 1951: 885 (cat.).
Krombein, 1979: 1508 (cat.).
MacLachlan, 1980, 53: 618.
Ahlstrom, 1995, 1: 102 (list).
Eumenes verticalis neoboreus ; Bohart, 1951: 885 (cat.).
Krombein, 1979: 1508 (cat.).
MacLachlan, 1980, 53: 618.
Eumenes verticalis coloradensis ; Bohart, 1951: 885.
Krombein, 1958: 163;
Krombein, 1979: 1508 (cat.).
MacLachlan, 1980, 53: 618.
Ruiz C. et al., 1993, 88: 83, 87 (list).
Rodríguez-Palafox, 1996: 479 (list).
Eumenes cruciferoides Giordani Soika, 1978 View in CoL , 29: 18 (key), 28. NEW SYNONYMY.
Type data: Holotype female at MSNV (examined).
Type locality: “ Messico: Nogares, dintorni di Orizaba, 1540 m .” (Mexico).
Rodríguez-Palafox, 1996: 479 (list).
Distribution: Canada (Alta, B.C., Queb, Man*, Ont., Sask*); Mexico (Lower California *, Nuevo León *, Tamaulipas *, Veracruz); U.S.A. (AZ, CA, CT, CO, DC*, DE*, ID, IL, IN*, MA, MD*, MN*, MO, NC, ND, NH, NJ, NM, NV*, NY, OH*, OR, PA*, SD*, UT, VA*, WI*, WV*, WY).
* Records from the literature.
Comments and diagnosis. Eumenes verticalis was originally described by Say (1824), from Pennsylvania (USA) (the type was destroyed), and currently it has four recognized subspecies: E. verticalis coloradensis Cresson, 1875 ; E. verticalis tricinctus Isely, 1917 , E. verticalis neoboreus Bequaert, 1944 , and E. verticalis verticalis (Say), 1824 .
We had examined all the type specimens from all the subspecies of E. verticalis plus 137 specimens (57 females and 80 males, all with male genitalia extracted and examined) and as we stated for E. consobrinus , E. verticalis has some intraspecific variation along its distribution, mainly in color and punctation of T2 throughout the east coast of the USA and southeast Canada, which leads us to propose the abandonment of these subspecies.
Because the primary type of the species was destroyed, we compared specimens of the typical E. verticalis with the original description of Say (1824) and we selected one of these for the redescription of the species. This specimen is established as the lectotype.
Giordani Soika (1978) described E. cruciferoides ( Fig. 12 View FIGURES 1–12 ) and noted that it resembles E. crucifera bolliformis but also noted that they differ in the “punctation of the clypeus, punctation and color (p. 29)”. We compared the type of E. cruciferoides and to the type of E. bolliformis and, although the former is known only from the type, we maintain it as a different species based on the following features: the punctures on clypeus are coarser and extend to the apical region; the punctation on the mesepisternum is denser, with the interspaces forming carinae, particularly in the center-basal region; the pubescence on the head and mesosoma is slightly longer, most evident on the frons and scape.
Although E. cruciferoides is known only from the type specimen, we compared it with the typical E. verticalis and no morphological differences were found besides having the ferrugineous marks contiguous with the yellow marks along body. As we observed for E. fraternus and E. crucifera var. bolliformis , specimens found the the southern USA and/or Mexico present ferrugineous marks along the body. As with E. verticalis , E. cruciferoides also has the clypeus with coarse punctation and long pubescence along the body, including on the scape (longer than the pedicel). For these reasons, we treat E. cruciferoides as a new synonym of E. verticalis .
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.
Kingdom |
|
Phylum |
|
Class |
|
Order |
|
Family |
|
Genus |
Eumenes verticalis Say, 1824
Grandinete, Yuri Campanholo, Noll, Fernando Barbosa & Carpenter, James 2018 |
Eumenes cruciferoides
Giordani Soika 1978 |
neoboreus
Bequaert 1944 |
Eumenes stenogaster
Isely 1917 |
Eumenes tricinctus
Isely 1917 |
Eumenes verticalis
Say 1824 |