Culicoides myanmaricus Szadziewski

Szadziewski, Ryszard, Dominiak, Patrycja, Sontag, Elżbieta, Krzemiński, Wiesław, Wang, Bo & Szwedo, Jacek, 2019, Haematophagous biting midges of the extant genus Culicoides Latreille (Diptera: Ceratopogonidae) evolved during the mid-Cretaceous, Zootaxa 4688 (4), pp. 535-548 : 541-542

Culicoides myanmaricus Szadziewski & Dominiak, sp. nov.

Figs 6 A–D

urn:lsid:zoobank.org:act:4671B5EB-F1B5-4F9C-830D-40E4965AA00A

Diagnosis. This species is distinguished from other Cretaceous Culicoides by the following combination of female characters: antennal flagellomere 1 with sensilla coeloconica, flagellomeres 2–8 subcylindrical to cylindrical, gradually increasing in length distally; palpal segment 3 slender, greatly elongate, without sensory pit; proboscis moderately elongate; wing membrane with macrotrichia on distal 1/3. Male unknown.

Description. Female ( Fig. 6A ). Head-body length minus antenna 1.15 mm. Proboscis ca. 0.25 mm long, 1.25 x longer than eye height. Eyes broadly separated ( Fig. 6B ). Antennal flagellum ( Fig. 6C ) ca. 0.39 mm long; first flagellomere with sensilla coeloconica; flagellomeres 2–13 gradually increasing in length; antennal ratio ca. 1.08. Palpus ( Fig. 6D ) 5-segmented, very slender, greatly elongate; total length 0.21 mm; segment 3 greatly elongate, without sensory pit, length 0.10 mm. Wing ( Fig. 6A ) length 0.63 mm; membrane with macrotrichia only on distal 1/3; basal radial cell without macrotrichia; costal ratio 0.81. Tarsomere 1 of hind leg slender; TR (1) 2.1, TR (3) 1.7. Abdominal sternite 8 with distinct caudomedian excavation.

Male. Unknown.

Material examined. Holotype female, NIGP 171165 (BA02 881). Burmese amber, Hukawng Valley , Myanmar. Syninclusion Hemiptera : Fulgoroidea of 3 rd or 4 th instar. Deposited in the collections of the Nanjing Institute of Ge- ology and Palaeontology, Chinese Academy of Sciences, Nanjing.

Etymology. The species name refers to Myanmar ( Burma) in Southeast Asia where Burmese amber is mined.

Figures

FIGURE 6. Fossil Culicoides (Groganomyia) myanmaricus Szadziewski & Dominiak, sp. nov., female holotype. A—total habitus, B—head, C—flagellum, D—palpus.