Inoderbe Shcherbakov et Emeljanov, 2021

Luo, Cihang, Song, Zhishun, Liu, Xiaojing, Jiang, Tian, Jarzembowski, Edmund A. & Szwedo, Jacek, 2022, Ingensalinae subfam. nov. (Hemiptera: Fulgoromorpha: Fulgoroidea: Inoderbidae), a new planthopper subfamily from mid-Cretaceous Kachin amber from Myanmar, Fossil Record 24 (2), pp. 455-465 : 456-457

Inoderbe Shcherbakov et Emeljanov, 2021 ; by original designation and monotypy.

Revised diagnosis. Rather small planthoppers, with habitus variable (derbid-like and tropiduchid-like). Lateral carinae of vertex and frons foliaceous, vertex and frons without median carina; fastigium may be developed; clypeus protruded, without median carina but with two lateral carinae. Pronotum with disc elevated, without median carina; postocular carinae may be developed; mesonotum in shape of diamond, with weakened or without distinct median and lateral carinae, mesoscutellum not distinctly separated. Tegmen with clear submarginal pterostigma, costal area wide with several transverse veinlets, not continued with apical cell series; clavus closed; stem MP forked much later than stem CuA with a few terminal branches (three), stem MP area reduced; stem CuA forked very early (at same level of fork of ScP + R or even earlier), with numerous branches (about six terminals), stem CuA area large, few crossveins in one gradate series. Hind wing with stems ScP + R, MP and CuA with a few branches (two or three), costal margin proximally with series of hamuli. Legs long and slender, carinate, metatibia without lateral spine, apical teeth without seta. Female ovipositor short, with three pairs of appendages: gonapophyses VIII, gonapophyses IX and gonoplac well distinguished, anal tube long, with arched side lobes; male pygofer tubular, slightly longer than wide, genital styles fused, male anal tube subquadrate and long, exceeding apices of genital styles.

Key to subfamilies of Inoderbidae

1. Body and appendages covered with filamentous wax, dorsum with numerous wax plates (in females only?) producing copious wax strands, especially long on posterior abdomen, wings not folded over abdomen in repose but held laterally, head narrow, frons with small fastigium at midline, antennae with scape and pedicel much elongate, tegmen slightly longer than body, narrowed to base, transparent, basal cell subtrapezoidal and much elongated, costal area with fewer than 10 transverse veinlets, CuA 1 with two terminal branches, clavus closed with acute apex, tornus very short, hind wing much shorter than tegmen, legs covered with numerous long setae. Inoderbinae Shcherbakov et Emeljanov, 2021, stat. nov.

2. Body and appendages not covered with filamentous wax, wings folded tectiform over abdomen in repose, head wide, frons without fastigium, antennae with scape and pedicel slightly elongate, tegmen much longer than body, very large and broad, translucent, basal cell relatively short, triangular, costal area with more than 10 transverse veinlets, CuA 1 single, clavus closed with blunt apex, tornus longer, hind wing slightly shorter than tegmen, legs covered with numerous very short setae. Ingensalinae Luo, Song et Szwedo subfam. nov.