Goja korytkowskii, Carrejo & Obando & Aldrete & Mendivil, 2021

Carrejo, Nancy, Obando, Ranulfo González, Aldrete, Alfonso Neri García & Mendivil, Julián, 2021, New species of Goja Navás (Psocodea: ‘ Psocoptera’: Epipsocidae) from Colombia Mexico and Peru, Zootaxa 4903 (4), pp. 501-541 : 521-524

Goja korytkowskii n. sp.

( Figs 56–66 )

Diagnosis. Male forewing with a vein from the lower angle of the pterostigma to the wing margin, a character so far unique in the genus; R2+3 unbranched, R4+5 forked. M five-branched, M5 forked. Hindwing Rs two branched, M two branched. Mesal endophallic sclerites large, broadly kidney-shaped; posterior endophallic sclerites with two slender, acuminate processes directed outward ( Fig. 61 ). Subgenital plate with sides converging to rounded apex. Ninth sternum broadly oval, wider anteriorly, with spermapore near posterior border. It is similar to G. horquetensis n. sp., G. farallones n. sp., G. caucana n. sp., G. munchinquensis n. sp., G. galarzai n. sp., G. circumvittata Casasola González & García Aldrete and G. tamensis n. sp. It differs from them in wing vention, phallosome and endophallic sclerites.

Male. Color (in 80% ethanol). Body pale brown. Head ( Fig. 58 ), brown; labral sclerites dark brown. Antennae brown. Compound eyes dark brown, ocelli hyaline, with ochre centripetal crescents. Maxillary palps brown. Antennae pale brown. Thorax brown. Legs brown. Forewings ( Fig. 56 ) brown, veins dark brown. Hindwings ( Fig. 57 ) light brown, veins pale brown.Abdomen pale brown, clunium brown. Hypandrium brown, dark brown on sides. Phallosome with aedeagal arch dark brown, endophallic sclerites dark brown; external parameres pale brown proximally, hyaline distally, internal margin dark brown. Epiproct and paraprocts brown.

Morphology. As in diagnosis, plus the following: Head: H/MxW: 1.38; compound eyes large, H/d: 4.47; IO/ MxW: 0.78. Outer cusp of lacinial tips broad, with nine denticles. Mx4/Mx2: 1.13. Forewings ( Fig. 56 ): FW/W: 2.68; pterostigma broadly triangular, wider in the middle: lp/wp: 3.66; vein Rs two-four branched, some specimens with accessory vein from the pterostigma to wing margin; areola postica wide, slanted posteriorly: al/ah: 2.82. Hindwings ( Fig. 57 ): HW/w: 3.09. Hypandrium ( Fig. 60 ) membranous, a field of distal macrosetae on each side of the longitudinal midline. Phallosome ( Fig. 61 ) slightly concave anteriorly; side struts straight, wide based and curved inwards; aedeagal arch with triangular posterior process; external parameres well developed, membranous, broad basally, apex narrow and rounded, with large pores. Paraprocts ( Fig. 59 ), ovoid, long, with setae and distal microsetae, large sensory fields with 30–31 trichobothria on basal rosettes. Epiproct ( Fig. 59 ) trapezoidal, wider basally, one macroseta mesally, two lateral macrosetae and small setae on distal field as illustrated.

Measurements. FW: 3525, HW: 2700, F: 850, T: 1400, t1: 580, t2: 190, ctt1: 23, f1: 630, f2: 450, Mx4: 210, IO: 405, D: 220, d: 160, IO/d: 2.53, PO: 0.73.

Female. Color (in 80% ethanol). As in the male, plus the following: forewing dark brown ( Fig. 62 ). Subgenital plate brown. Gonapophyses brown. IX sternum yellowish.

Morphology. Head: H/MxW: 1.48; compound eyes large, H/d: 4.67; IO/MxW: 0.76. Ocelli absent; outer cusp of lacinial tips broad, with eight-nine denticles. Mx4/Mx2: 1.03. Forewings reduced to small scales, ovoid, setose, without defined veins ( Fig. 62 ): FW/W: 1.05. Hindwings absent. Legs: tarsi without ctenidobotria. Subgenital plate ( Fig. 65 ) triangular, broad, posteriorly rounded. Gonapophyses ( Fig. 66 ): v1 elongate, narrow basally and wider subapically, acuminate distally, surpassing v3; v2+3 triangular, with 8–9 setae on v3; distal process long, straight, acuminate, with field of microspines; IX sternum ( Fig. 66 ) oval, spermapore posteriorly. Paraprocts ( Fig. 64 ), triangular, wide, with distal setal field, large sensory fields with 33 trichobothria on basal rosettes. Epiproct ( Fig. 64 ) trapeziform, posterior margin with setal field, one macroseta mesally.

Measurements. FW: 475, HW: 700, T: 1025, t1: 450, t2: 180, ctt1: 0.0, f1: 340, f2: 280, f3: 230, Mx4: 200, IO: 397, D: 225, d: 165, IO/d: 2.41, PO: 0.73.

Specimens studied. Holotype male. COLOMBIA. Valle del Cauca, Santiago de Cali, Felidia, El Pato , ca. La Teresita , 3°26’34.6”N: 76°39’46.7”W, 2058 m., 14.X.2011, R. González & L. Villegas. Beating vegetation and dead branches . Paratypes: 2 males, 2 females, Santiago de Cali , K 18, ca. Zíngara, 3°30’38.3”N: 76°37’13.8”W, 1990 m., 6.IV.2012, R. González & N. Carrejo . 2 males, 1 female, El Cerrito , Tenerife , Natural Reserve El Pailón, 3°42’16.7”N: 76°03’54.3”W, 2235 m., 5.VII.2019, R. González. Beating vegetation and dead branches .

Etymology. This species is dedicated, in memoriam, to Cheslavo Korytkowski, dipterologist, specialized in the neotropical dipterofauna.

Figures

FIGURES 56‒61. G. korikowskii n. sp. Male. 56. Forewing. 57. Hindwing. 58. Front view of head. 59. Left paraproct and epiproct. 60. Hypandrium. 61. Phallosome. Scales in mm.

FIGURES 62‒66. G. korytkowskii n. sp. Female. 62. Forewing. 63. Front view of head. 64. Left paraproct and epiproct. 65. Subgenital plate. 66. Ninth sternum and left gonapophyses. Scales in mm.