Camponotus christi Forel

Rakotonirina, Jean Claude & Fisher, Brian L., 2018, Taxonomic revision of the Malagasy Camponotus subgenus Mayria (Hymenoptera, Formicidae) using qualitative and quantitative morphology, Zootaxa 4438 (1), pp. 1-58 : 19-21

publication ID	https://doi.org/ 10.11646/zootaxa.4438.1.1
publication LSID	lsid:zoobank.org:pub:55BEB1B1-DA14-4CC4-92EB-1EA98AD6277C
DOI	https://doi.org/10.5281/zenodo.5987962
persistent identifier	https://treatment.plazi.org/id/03C287BB-E879-FF9D-31A6-4E51FBBEF87D
treatment provided by	Plazi
scientific name	Camponotus christi Forel
status

Treatment

Camponotus christi Forel View in CoL

( Figures 8B View FIGURE 8 , 9B View FIGURE 9 , 12A View FIGURE 12 , 13A View FIGURE 13 , 15 View FIGURE 15 , 21 View FIGURES 21–26 )

Camponotus christi Forel, 1886b: 184 View in CoL . Lectotype minor worker, present designation, Centre de Madagascar [Ambatomanjaka, Miarinarivo, -18.766947, 46.869107, 1344 m] (Hildebrandt), AntWeb CASENT0101544 (MHNG). Paralectotype: 1 minor worker with same data as lectotype but specimen coded: CASENT0101435 (Camboué) (MNHN) [examined]. [Combination in Camponotus (Myrmocamelus) View in CoL : Forel, 1914: 270; in Camponotus (Myrmosaga) View in CoL : Wheeler, 1922: 1045; in Camponotus (Mayria) View in CoL : Emery, 1925: 122; Bolton, 1995: 92, 131].

Diagnosis. Median portion of clypeus without longitudinal carina; mandible with six teeth; anteromedian margin of clypeus not bordered by a lamella; in lateral view, mesosoma long and low, its dorsal outline not a dome-like

structure; propodeal declivity inclined anteriorly; body color entirely yellow to light brown, legs with same color as body.

Description. Minor worker. In full-face view, head elongate (CWb/CL: 0.76±0.03; 0.69–0.83), lateral margins straight and more or less parallel to each other, rounding broadly to the slightly, medially excised posterior

margin. Anterior clypeal margin straight, with broadly angulate junction to lateral margin; median longitudinal carina absent. Eyes not breaking lateral outlines of head; their posterior margin located approximately at posterior third portion of head (PoOc/CL: 0.34±0.01; 0.31–0.36). Mandible triangular, apical margin with six sharp teeth. Antennal scape long, more than its distal half extending beyond posterior cephalic border. In lateral view, pronotum and anterior half of mesonotum forming feeble convexity, posterior half of mesonotum and propodeal dorsum straight and long, joining declivity surface with noticeable angle. Propodeal dorsum roughly three times as long as height of declivity; propodeal spiracle circular. Petiolar node as high as or higher than long, its dorsal margin rounding to anterior face and joining posterior face at an angle.

Dorsum of head covered with numerous yellowish, elongate, erect hairs; pronotum and mesonotum each with a pair of erect hairs, propodeum with a pair of hairs found on junction to declivity; sparse, short pubescence present on dorsum of body. Body color entirely yellowish-orange to orange-brown and funicular segments becoming dark brown toward the apex.

Major worker. Differing from minor worker in the following characters: enlarged head, with markedly concave posterior margin; apical third of antennal scape surpassing posterior cephalic margin; robust mesosoma, metanotum distinctly visible, propodeal dorsum rounding to declivity and twice as long as height of declivity; petiolar node tapering dorsally. More pairs of erect hairs on junction of propodeal dorsum and declivity and on posterodorsal margin of petiolar node.

Discussion. Workers of C. christi are similar to those of C. mainty , C. foersteri and C. maculiventris in that their clypeus lacks a median longitudinal carina, and in lateral view, their mesosoma is long and low. However, C. christi workers are easily separable by their entirely yellow to light brown body color and legs. In the members of C. mainty , their body is entirely black to reddish-black. In the two latter species, at least the trochanters and the distal portion of the coxa are yellowish to white.

The cluster of C. christi shown by the dendrogram of NC-PART clustering is confirmed by the cumulative LDA at 100% identification success, indicating the support of the species hypothesized by qualitative morphologybased taxonomy. The cluster for C. christi is located next to that of C. foersteri and C. mainty , which suggests that the three species are more similar to each other than other species. This finding corroborates their close morphological similarity as detected by the qualitative analysis.

Distribution and biology. Camponotus christi is widely distributed across Madagascar. It occupies sclerophyll forests of the central high plateau, tropical dry forests in the west to coastal scrub and littoral rainforests, as well as montane rainforests in the east. While its workers have been found foraging on the forest floor and the lower portions of vegetation, their nest sites have been located in root mats in the ground layer, rotten logs, rotting tree stumps, rotten pockets above the ground, dead branches and bamboo above the ground, and in canopy moss and leaf litter.

Additional material examined. ‘Andrangoloaka], (Sikora) (MHNG); [antananarivo, Museum Paris, Grandidier 1893], (Camboué) (MNHN); [Divers, tous les Divers sont de la Forêt d’Andrangoloaka, 20, Museum Paris, Grandidier 1899] (MNHN); [ Forêt d’Analamanita Province Antananarivo], (Camboué) ( MHNG) ; 3 km 41° NE Andranomay, 11.5 km 147° SSE Anjozorobe, 1300 m, -18.47333, 47.96, montane rainforest, (Fisher, Griswold et al.) (CASC); Analamanga Region, District of Ankazobe, Ambohitantely , 46 km NE of Ankazobe, 701 m, - 18.198, 47.2815, sclerophyll forest, (Mike, Rinha) ( CASC) ; Forêt de galerie, Andranorovitra, 24.0 km NNE Ankazobe , 1491 m, -18.11243, 47.19757, disturbed gallery montane forest, (B.L. Fisher et al.) ( CASC) ; Mandraka , 1312 m, -18.91813, 47.91717, montane rainforest, (B.L. Fisher et al.) ( CASC) ; Mandraka Park , 1360 m, -18.9019, 47.90786, montane shrubland, (B.L. Fisher et al.) ( CASC) ; Réserve Spéciale d'Ambohitantely, Forêt d Ambohitantely, 20.9 km 72° NE d Ankazobe , 1410 m, -18.22528, 47.28683, montane rainforest, (Fisher, Griswold et al.) ( CASC) ; Réserve Spéciale d'Ambohitantely , 1580 m, -18.18762, 47.28576, montane forest, (B.L. Fisher et al.) ( CASC) ; Réserve Spéciale d'Ambohitantely , 1490 m, -18.22444, 47.2774, montane forest, (B.L. Fisher et al.) ( CASC) ; Tananarive, (Waterlot) (MNHN); Province Antsiranana: [Diego Suarez], (Ch. Alluaud) ( MHNG) ; 11.0 km WSW Befingotra, Réserve Anjanaharibe-Sud, 1565 m, -14.75, 49.45, montane rainforest, (B.L. Fisher) (CASC); 6.5 km SSW Befingotra, Réserve Anjanaharibe-Sud , 875 m, -14.75, 49.5, rainforest, (B.L. Fisher) ( CASC) ; 9.2 km WSW Befingotra, Réserve Anjanaharibe-Sud , 1280 m, -14.75, 49.46667, montane rainforest, (B.L. Fisher) ( CASC) ; 9.2 km WSW Befingotra, Réserve Anjanaharibe-Sud , 1200 m, -14.75, 49.46667, montane rainforest, (B.L. Fisher) ( CASC) ; Ampasindava, Andranomatavy Forest , 543 m, -13.66296, 47.97936, disturbed dry forest, (B.L. Fisher et al.) ( CASC) ; Ampasindava, Forêt d'Ambilanivy, 3.9 km 181° S Ambaliha , 600 m, - 13.79861, 48.16167, rainforest, (Fisher, Griswold et al.) ( CASC) ; Antsiranana II Pref: Antsahampano S. - Pref: Montagne d'Ambre. Site MD1, 1049 m, -12.52765, 49.17235, in Commelina regrowth on path next to degraded primary riparian rainforest, (D. Lees, R. Ranaivosolo & P. Razafindraibe) ( CASC) ; Betaolana Forest, along Bekona River , 880 m, -14.52996, 49.44039, rainforest, (B.L. Fisher et al.) ( CASC) ; Forêt de Binara , 9.1 km 233° SW Daraina, 725 m, -13.26333, 49.60333, rainforest, (B.L. Fisher) ( CASC) ; Galoko chain, Mont Galoko , 520 m, - 13.58487, 48.71818, rainforest, (B.L. Fisher et al.) ( CASC) ; Galoko chain, Mont Galoko , 980 m, -13.5888, 48.72864, montane forest, (B.L. Fisher et al.) ( CASC) ; Galoko chain, Mont Galoko , 1100 m, -13.59358, 48.73157, montane forest, (B.L. Fisher et al.) ( CASC) ; Makirovana forest , 415 m, -14.17066, 49.95409, rainforest, (B.L. Fisher et al.) ( CASC) ; Makirovana forest , 715 m, -14.16666, 49.95, rainforest, (B.L. Fisher et al.) ( CASC) ; Makirovana forest , 390 m, -14.10295, 50.01984, rainforest, (B.L. Fisher et al.) ( CASC) ; Makirovana forest , 715 m, -14.16666, 49.95, rainforest, (B.L. Fisher et al.) ( CASC) ; Montagne d'Ambre National Park, Roussette Camp 7 km SW Park entrance, 960 m, -12.51444, 49.18139, rainforest, (Mike, Frank, Rin'ha) ( CASC) ; Parc National de Marojejy, 25.7 km 32° NNE Andapa, 10.3 km 314° NW Manantenina , 1575 m, -14.445, 49.74167, montane rainforest, (B.L. Fisher) ( CASC) ; Parc National de Marojejy, Antranohofa , 26.6 km 31° NNE Andapa, 10.7 km 318° NW Manantenina, 1325 m, -14.44333, 49.74333, montane rainforest, (B.L. Fisher) ( CASC) ; Parc National de Marojejy, Manantenina River , 27.6 km 35° NE Andapa, 9.6 km 327° NNW Manantenina, 775 m, -14.435, 49.76, rainforest, (B.L. Fisher et al.) ( CASC) ; Parc National Montagne d'Ambre , 1100 m, -12.52306, 49.17901, montane rainforest, (B.L. Fisher et al.) ( CASC) ; Parc National Montagne d'Ambre , 3.6 km 235° SW Joffreville, 925 m, - 12.53444, 49.1795, montane rainforest, (Fisher, Griswold et al.) ( CASC) ; Parc National Montagne d'Ambre, Mahasarika, 1135 m, -12.53176, 49.17662, montane rainforest, (B.L. Fisher et al.) (CASC); Parc National Montagne d'Ambre, Roussettes, 1025 m, -12.52574, 49.17238, montane rainforest, (B.L. Fisher et al.) (CASC); Réserve Spéciale Manongarivo , 12.8 km 228° SW Antanambao, 780 m, -13.97667, 48.42333, rainforest, (B.L. Fisher) ( CASC) ; SAVA Region, District of Sambava, Marojejy National Park, 5 km W of Manantenina village, 1st Camp site ( Mantella ), 487 m, -14.43817, 49.774, Low altitude rainforest, (Rin'ha, Mike) ( CASC) ; 3 km W Sakalava Beach [white dunes site], 40 m, -12.28617, 49.36667, white dunes in littoral forest, (Harin'Hala, Irwin, Schlinger) ( CASC) ; Montaigne Français , 150 m, -12.325, 49.33333, along forested limestone ridge, (R. Harin'Hala) ( CASC) ; Parc National Montagne d'Ambre [1 st campsite ], 960 m, -12.51444, 49.18139, rainforest, (R. Harin'Hala) ( CASC) ; Parc National Montagne d'Ambre [ Petit Lac road], 1125 m, -12.52028, 49.17917, rainforest, (R. Harin'Hala) ( CASC) ; Sakalava Beach [vegetated beach dunes], 10 m, -12.26278, 49.3975, across sandy trail in dwarf littoral forest, (R. Harin'Hala) ( CASC) ; Province Fianarantsoa: [Imerina], (Camboué) ( MHNG) ; [Nosi be Imerina], (Sikora) ( MHNG); 3 km W Ranomafana , nr . Ifandiana , 950 m, -21.25, 47.41667, rainforest, (P.S. Ward) ( PSWC) ; 38 km S Ambalavao, Réserve Andringitra , 1680 m, -22.2, 46.96667, montane rainforest, (B.L. Fisher) ( CASC) ; 40 km S Ambalavao, Réserve Andringitra , 1275 m, -22.21667, 46.96667, montane rainforest, (B.L. Fisher) ( CASC) ; 7.6 km 122° Kianjavato, Forêt Classée Vatovavy, 175 m, -21.4, 47.94, rainforest, (B.L. Fisher et al.) (CASC); 8.0 km NE Ivohibe, 1200 m, -22.42167, 46.89833, montane rainforest, (B.L. Fisher & Sylvain) (CASC); 9.0 km NE Ivohibe, 900 m, -22.42667, 46.93833, rainforest, (B.L. Fisher & Sylvain) (CASC); Belle Vue trail, Ranomafana National Park , Fianarantsoa Prov., 1020 m, -21.2665, 47.42017, mixed tropical forest, (M.E. Irwin, F.D. Parker, R. Harin'Hala) ( CASC) ; Fitovavy Fitovinany Region, District of Ifanadiana Belle vue area 1200 m S of Ranomafana National Park entrance, 1018 m, -21.2665, 47.42017, rainforest, (Rin'ha, Mike) ( CASC) ; Fitovavy Fitovinany Region, District of Ifanadiana , 12 km W of Ranomafana, 1127 m, -21.25083, 47.40717, forest edge, open area, (Rin'ha, Mike) ( CASC) ; Forêt d'Atsirakambiaty , 7.6 km 285° WNW Itremo, 1550 m, -20.59333, 46.56333, montane rainforest, (Fisher, Griswold et al.) ( CASC) ; Forêt de Vevembe, 66.6 km 293° Farafangana , 600 m, -22.791, 47.18183, rainforest, transition to montane forest, (B.L. Fisher et al.) ( CASC) ; JIRAMA water works near river, Ranomafana National Park , Fianarantsoa Prov., 690 m, -21.2485, 47.45217, open area near stream, (M.E. Irwin, F.D. Parker, R. Harin'Hala) ( CASC) ; Réserve Spéciale Manombo , 32 km SE of Farafangana, 36 m, -23.02183, 47.72, Lowland rainforest, (Rin'ha, Mike) ( CASC) ; Miandritsara Forest , 40 km S of Ambositra, 822 m, -20.79267, 47.17567, Low altitude rainforest, (Rin'ha, Mike) ( CASC) ; Parc National Befotaka-Midongy, Papango 27.7 km S Midongy-Sud, Mount Papango , 940 m, -23.83517, 46.96367, rainforest, (B.L. Fisher et al.) ( CASC) ; Parc National Befotaka-Midongy, Papango 28.5 km S Midongy-Sud, Mount Papango , 1250 m, - 23.84083, 46.9575, montane rainforest, (B.L. Fisher et al.) ( CASC) ; Parc National de Ranomafana, Vatoharanana River , 4.1 km 231° SW Ranomafana, 1100 m, -21.29, 47.43333, montane rainforest, (Fisher, Griswold et al.) ( CASC) ; Réserve Spéciale Ivohibe 8.0 km E Ivohibe, 1200 m, -22.48333, 46.96833, montane rainforest, (B.L. Fisher & Sylvain) ( CASC) ; Réserve Spéciale Ivohibe , 7.5 km ENE Ivohibe, 900 m, -22.47, 46.96, rainforest, (B.L. Fisher & Sylvain) ( CASC) ; Réserve Spéciale Manombo 24.5 km 228° Farafangana , 30 m, -23.01583, 47.719, rainforest, (B.L. Fisher et al.) ( CASC) ; radio tower, Ranomafana National Park , Fianarantsoa Prov., 1130 m, - 21.25083, 47.40717, forest edge, mixed tropical forest, open area, (M. Irwin, R. Harin'Hala) ( CASC) ; Ranomafana National Park, Talatakely area , 0.4 km WSW of Park Entrance, 900 m, -21.41667, 47.68333, mixed tropical forest, (D. H. & K.M. Kavanaugh) ( CASC) ; stream area, 900 m E of Isalo National Park Interpretive Center, Fianarantsoa Prov ., 750 m, -22.62667, 45.35817, open area near stream, (R. Harin'Hala) (CASC); Vohiparara, -21.23333, 47.36667, (A. Pauly) (CASC); Vohiparara broken bridge, Fianarantsoa Prov., 1110 m, -21.22617, 47.36983, high altitude rainforest, (R. Harin'Hala) ( CASC) ; Province Mahajanga: Forêt de Tsimembo , 8.7 km 336° NNW Soatana, 20 m, -19.02139, 44.44067, tropical dry forest, (Fisher-Griswold Arthropod Team) ( CASC) ; Réserve Spéciale Marotandrano, Marotandrano 48.3 km S Mandritsara , 865 m, -16.28322, 48.81443, transition humid forest, (B.L. Fisher et al.) ( CASC) ; Region Sofia, Bemanevika, 1606 m, -14.337, 48.58874, montane rainforest, (B.L. Fisher et al.) (CASC); Region Sofia, Bemanevika, 1657 m, -14.32826, 48.58406, montane rainforest, (B.L. Fisher et al.) (CASC); Province Toamasina: Baie d’Antongil (Mocquerys) ( MSNG) ; 19 km ESE Maroantsetra , 250 m, -15.48333, 49.9, rainforest, (P.S. Ward) ( PSWC) ; 1 km SSW Andasibe (= Perinet ), 920 m, -18.93333, 48.41667, rainforest, (P.S. Ward) ( PSWC) ; 5.3 km SSE Ambanizana, Andranobe , 600 m, -15.66667, 49.96667, rainforest, (B.L. Fisher) ( CASC) ; 5.3 km SSE Ambanizana, Andranobe , 425 m, -15.67133, 49.97395, rainforest, (B.L. Fisher) ( CASC) ; 6.3 km S Ambanizana, Andranobe , 25 m, -15.6813, 49.958, rainforest, (B.L. Fisher) ( CASC) ; 6.9 km NE Ambanizana, Ambohitsitondroina , 825 m, -15.58506, 50.00952, rainforest, (B.L. Fisher) ( CASC) ; 6.9 km NE Ambanizana, Ambohitsitondroina , 1000 m, -15.56667, 50, montane rainforest, (B.L. Fisher) ( CASC) ; 6.9 km NE Ambanizana, Ambohitsitondroina , 1080 m, -15.56667, 50, montane rainforest, (B.L. Fisher) ( CASC) ; 6 km ESE Andasibe (= Perinet ), 900 m, -18.95, 48.46667, rainforest, (P.S. Ward) ( PSWC) ; 7 SE Andasibe National Park Headquarters , 1050 m, -18.96278, 48.45267, tropical forest, (M.E. Irwin, R. Harin'Hala) ( CASC) ; Ambalahasina , 62.4 km 19° Toamasina, 15 m, -17.59452, 49.46785, coastal scrub, (B. Blaimer, F.N. Raharimalala) ( CASC) ; Ambatovy , 12.4 km NE Moramanga, 1010 m, -18.84963, 48.2947, montane rainforest, (B.L. Fisher et al.) ( CASC) ; Ambatovy , 12.4 km NE Moramanga, 1000 m, -18.84773, 48.29568, montane rainforest, (B.L. Fisher et al.) ( CASC) ; Ambatovy , 12.4 km NE Moramanga, 1080 m, -18.83937, 48.30842, montane rainforest, (B.L. Fisher et al.) ( CASC) ; Analamay , 1068 m, -18.80623, 48.33707, montane rainforest, (Malagasy ant team) ( CASC) ; Andasibe National Park , botanic garden near entrance, west of ANGAP office, 1025 m, -18.92639, 48.40783, tropical forest, (M.E. Irwin, R. Harin'Hala) ( CASC) ; Ankerana , 865 m, -18.40062, 48.81311, rainforest, (B.L. Fisher et al.) ( CASC) ; Ankerana , 750 m, -18.40829, 48.82107, rainforest, (B.L. Fisher et al.) ( CASC) ; Ankerana , 1035 m, -18.4017, 48.80605, montane forest, (B.L. Fisher et al.) ( CASC) ; Bevolota 17.1 km N Andasibe, 995 m, - 18.77071, 48.43164, montane rainforest, (B.L. Fisher et al.) ( CASC) ; Corridor Forestier Analamay-Mantadia, Ambatoharanana, 1016 m, -18.79944, 48.40375, rainforest, (B.L. Fisher et al.) (CASC); Corridor Forestier Analamay-Mantadia, Ambatoharanana, 968 m, -18.80424, 48.40081, rainforest, (B.L. Fisher et al.) (CASC); Corridor Forestier Analamay-Mantadia, Ambatoharanana, 1013 m, -18.80388, 48.40506, rainforest, (B.L. Fisher et al.) (CASC); Corridor Forestier Analamay-Mantadia, Ambatoharanana, 968 m, -18.80424, 48.40081, rainforest, (B.L. Fisher et al.) (CASC); Corridor Forestier Analamay-Mantadia, Ambatoharanana, 1064 m, -18.80398, 48.40358, rainforest, (B.L. Fisher et al.) (CASC); Corridor Forestier Analamay-Mantadia, Ambatoharanana, 968 m, -18.80424, 48.40081, rainforest, (B.L. Fisher et al.) (CASC); Corridor Forestier Analamay-Mantadia, Ambatoharanana, 960 m, -18.80438, 48.40735, rainforest, (B.L. Fisher et al.) (CASC); Corridor Forestier Analamay-Mantadia, Ambohibolakely, 1044 m, -18.76087, 48.37128, rainforest, (B.L. Fisher et al.) (CASC); Corridor Forestier Analamay-Mantadia, Ambohibolakely, 983 m, -18.76131, 48.36437, rainforest, (B.L. Fisher et al.) (CASC); Corridor Forestier Analamay-Mantadia, Ambohibolakely, 918 m, -18.77898, 48.36375, rainforest, (B.L. Fisher et al.) (CASC); Corridor Forestier Analamay-Mantadia, Ambohibolakely, 983 m, -18.76131, 48.36437, rainforest, (B.L. Fisher et al.) (CASC); Corridor Forestier Analamay-Mantadia, Tsaravoniana, 984 m, - 18.76369, 48.4203, rainforest, (B.L. Fisher et al.) (CASC); Corridor Forestier Analamay-Mantadia, Tsaravoniana, 1039 m, -18.76465, 48.41938, rainforest, (B.L. Fisher et al.) (CASC); Corridor Forestier Analamay-Mantadia, Tsaravoniana, 1018 m, -18.75737, 48.42302, rainforest, (B.L. Fisher et al.) (CASC); F.C. Andriantantely, 530 m, - 18.695, 48.81333, rainforest, (H.J. Ratsirarson) (CASC); F.C. Didy, 960 m, -18.19833, 48.57833, rainforest, (H.J. Ratsirarson) (CASC); F.C. Sandranantitra, 450 m, -18.04833, 49.09167, rainforest, (H.J. Ratsirarson) (CASC); Forêt Ambatovy, 14.3 km 57° Moramanga , 1075 m, -18.85083, 48.32, montane rainforest, (Malagasy ant team) ( CASC) ; Manakambahiny Atsinanana, -17.75, 48.71667, Primary forest , (A. Pauly) ( CASC) ; Montagne d'Akirindro 7.6 km 341° NNW Ambinanitelo, 600 m, -15.28833, 49.54833, rainforest, (Fisher, Griswold et al.) ( CASC) ; Montagne d'Anjanaharibe, 18.0 km 21° NNE Ambinanitelo , 470 m, -15.18833, 49.615, rainforest, (Fisher, Griswold et al.) ( CASC) ; Montagne d'Anjanaharibe , 19.5 km 27° NNE Ambinanitelo, 1100 m, -15.17833, 49.635, montane rainforest, (Fisher, Griswold et al.) ( CASC) ; Nosy Mangabe , 300 m, -15.5, 49.76667, rainforest, (P.S. Ward) ( PSWC) ; Nosy Mangabe , 5 m, -15.5, 49.76667, littoral vegetation, (P.S. Ward) ( PSWC) ; Nosy Mangabe , 7.43 km S Maroantsetra, 3 m, -15.4973, 49.76223, littoral rainforest, (B.L. Fisher et al.) ( CASC) ; Parc National de Masoala , 39.4 km 150° SSE Maroantsetra, 200 m, -15.71, 49.97, rainforest, (B.L. Fisher, H.J. Ratsirarson) ( CASC) ; Parc National Mananara-Nord, 7.1 km 261° Antanambe , 225 m, -16.455, 49.7875, rainforest, (B.L. Fisher et al.) ( CASC) ; Parcelle K 7 Tampolo , 10 m, -17.28333, 49.41667, littoral forest, (Malagasy ant team) ( CASC) ; Réserve Nationale Integrale Betampona, Betampona 35.1 km NW Toamasina , 500 m, - 17.91801, 49.20074, rainforest, (B.L. Fisher et al.) ( CASC) ; Réserve Spéciale Ambatovaky, Sandrangato river , 450 m, -16.77274, 49.26551, rainforest, (B.L. Fisher et al.) ( CASC) ; Réserve Spéciale Ambatovaky, Sandrangato river , 475 m, -16.76912, 49.26704, rainforest, (B.L. Fisher et al.) ( CASC) ; Réserve Spéciale Ambatovaky, Sandrangato river , 520 m, -16.7633, 49.26692, rainforest, (B.L. Fisher et al.) ( CASC) ; Réserve Spéciale Ambatovaky, Sandrangato river , 470 m, -16.7702, 49.26638, rainforest, (B.L. Fisher et al.) ( CASC) ; Réserve Spéciale Ambatovaky, Sandrangato river , 360 m, -16.81753, 49.29498, rainforest, (B.L. Fisher et al.) ( CASC) ; Réserve Spéciale Ambatovaky, Sandrangato river , 425 m, -16.8162, 49.29202, rainforest, (B.L. Fisher et al.) ( CASC) ; Réserve Spéciale Ambatovaky, Sandrangato river , 460 m, -16.81209, 49.29216, rainforest, (B.L. Fisher et al.) ( CASC) ; Réserve Spéciale Ambatovaky, Sandrangato river , 480 m, -16.80561, 49.29507, rainforest, (B.L. Fisher et al.) ( CASC) ; Réserve Ambodiriana, 4.8 km 306° Manompana, along Manompana river , 125 m, -16.67233, 49.70117, rainforest, (B.L. Fisher et al.) ( CASC) ; Réserve Perinet-Analamazaotra , 950 m, -18.93333, 48.43333, rainforest, (D. M. O lson) ( PSWC) ; Réserve Betampona, Camp Rendrirendry 34.1 km 332° Toamasina, 390 m, - 17.924, 49.19967, rainforest, (B.L. Fisher et al.) ( CASC) ; S.F. Tampolo, 10 km NNE Fenoarivo Atn. , 10 m, - 17.2825, 49.43, littoral rainforest, (B.L. Fisher) ( CASC) ; Tampolo, Masoala Peninsula, 40.4 km 154° SSE Maroantsetra , 30 m, -15.73, 49.96, rainforest, (B.L. Fisher, H.J. Ratsirarson) ( CASC) ; Torotorofotsy , 1005 m, - 18.77048, 48.43043, montane rainforest, (B.L. Fisher et al.) ( CASC) ; Torotorofotsy , 1070 m, -18.87082, 48.34737, montane rainforest, marsh edge, (Malagasy ant team) ( CASC) ; Province Toliara: 10 km NW Enakara, Res Andohahela , 420 m, -24.56667, 46.81667, rainforest, (B.L. Fisher) ( CASC) ; 13 km NW Enakara, Réserve Andohahela , 1250 m, -24.55, 46.8, montane rainforest, (B.L. Fisher) ( CASC) ; 13 km NW Enkara, Res Andohahela , 1140 m, -24.55, 46.8, montane rainforest, (B.L. Fisher) ( CASC) ; 6 km ESE Imonty, Réserve Andohahela , 1200 m, -24.85, 46.75, rainforest, (P.S. Ward) ( PSWC) ; Anosy Region, Anosyenne Mts , 29.9 km NW Manantenina, 750 m, -24.13894, 47.06804, montane rainforest, (B.L. Fisher, F.A. Esteves et al.) ( CASC) ; Anosy Region, Anosyenne Mts , 31.2 km NW Manantenina, 1125 m, -24.13894, 47.06804, rainforest, (B.L. Fisher, F.A. Esteves et al.) ( CASC) ; Anosy Region, Anosyenne Mts , 31.2 km NW Manantenina, 1315 m, -24.13632, 47.05485, rainforest, (B.L. Fisher, F.A. Esteves et al.) ( CASC) ; Anosy Region, Parc National Andohahela, Col de Tanatana , 400 m, -24.74969, 46.84949, rainforest, (B.L. Fisher, F.A. Esteves et al.) ( CASC) ; Forêt Ivohibe 55.0 km N Tolagnaro, 200 m, -24.569, 47.204, rainforest, (B.L. Fisher et al.) ( CASC) ; Forêt Ivohibe 55.6 km N Tolagnaro, 650 m, -24.56167, 47.20017, rainforest, (B.L. Fisher et al.) ( CASC) ; Parc National Andohahela, Col de Tanatana , 33.3 km NW Tolagnaro, 275 m, -24.7585, 46.85367, rainforest, (B.L. Fisher et al.) ( CASC) ; Parc National Andohahela, Manangotry, 33.8 km NW Tolagnaro , 575 m, -24.75117, 46.85783, rainforest, (B.L. Fisher et al.) ( CASC) ; Parc National d'Andohahela, Col du Sedro, 3.8 km 113° ESE Mahamavo, 37.6 km 341° NNW Tolagnaro , 900 m, -24.76389, 46.75167, montane rainforest, (Fisher-Griswold Arthropod Team) ( CASC) ; Réserve Privée Berenty, Forêt de Bealoka, Mandrare River , 14.6 km 329° NNW Amboasary, 35 m, -24.95694, 46.2715, gallery forest, (Fisher-Griswold Arthropod Team) ( CASC) ; Réserve Spéciale Kalambatritra , 1365 m, -23.4185, 46.4583, grassland, (B.L. Fisher et al.) ( CASC) ; Réserve Spéciale Kalambatritra, Ambinanitelo, 1325 m, -23.4502, 46.45658, montane rainforest, (B.L. Fisher et al.) (CASC); Réserve Spéciale Kalambatritra, Ampanihy, 1270 m, - 23.4635, 46.4631, montane rainforest, (B.L. Fisher et al.) (CASC); Réserve Spéciale Kalambatritra, Ampanihy, 1269 m, -23.463, 46.47057, montane rainforest, (B.L. Fisher et al.) (CASC); [ Central Madagascar ], ( MHNG) ; [ Central Madagascar ], ( MHNG) ; [de Diversa, Museum Paris, Grandidier 1899], (MNHN); [Madagasc.], (MHNG); 11 km SE Ampasimanolotra (= Brickaville ), 5 m, -18.9, 49.13333, littoral rainforest, (P.S. Ward) ( PSWC) ; massif du Tsaratanana (versant Sud), Andohanambatoafo , 2030 m, (P. Soga) ( MNHN) .