Philonthus molliculus, Casey, 1915

Herman, Taphylininae Iochini Aorothiini Thiini Latyprosopini Taphylinini Mblyopinina Nisolinina Yptiomina Hilonthina Lee H., 2001, Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the End of the Second Millennium. V. Staphylinine Group (Part 2) Staphylininae: Diochini, Maorothiini, Othiini, Platyprosopini, Staphylinini (Amblyopinina, Anisolinina, Hyptiomina, Philonthina), Bulletin of the American Museum of Natural History 2001 (265), pp. 2441-3020 : 2441-3020

molliculus Casey, 1915 , see: distans Horn, 1884 .

montanus Bernhauer, 1934 g: 237 ( Philonthus ; [preoccupied]; Type locality: Ruwenzori: Kalonge, 2050 m; Kivu: Mts. Nyamukubi , 2600 m).

— Tottenham, 1956b: 207 ( Gabrius ; characters).

— Herman, 2001: 44 ( Gabrius ; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).

- DISTRIBUTION: Congo.

monticola Bernhauer, 1931d: 586 ( Philonthus ; subgenus Gabrius ; Type locality: Abyssinia: Mt. Zukwala, ca. 9000 ft. and 9600 ft.; Mt. Chillalo, ca. 9000 ft.; Jem-Jem Forest,

ca. 9,000 ft.).

— Scheerpeltz, 1933: 1351 ( Philonthus ; catalog).

— Bernhauer and Chapman, 1939: 71 ( Philonthus ; Kenya).

— Tottenham, 1949b: 362 ( Gabrius ; characters).

— Tottenham, 1956: 305 ( Gabrius ; characters in key).

— Tottenham, 1956b: 206, 207, 211 ( Gabrius ; characters).

- DISTRIBUTION: Ethiopia, Kenya.

morator Schillhammer, 1997: 68 ( Gabrius ; Type locality: Indonesia: W-Sumatra, Singgalang , 2100–2800 m).

- DISTRIBUTION: Indonesia.

mubilibanus Cameron, 1950 g: 42 ( Philonthus ; subgenus Gabrius ; Type locality: Belgian Congo: Mubiliba, volc. Nyamuragira, alt. 2000 m).

— Schillhammer, 1997: 79 ( Gabrius ).

- DISTRIBUTION: Congo.

muelleri Gridelli, 1928: 47 ( Gabrius ; Type locality: Lenkoran, Caucaso ).

— Scheerpeltz, 1933: 1352 ( Philonthus ; catalog).

— Smetana, 1961: 37 ( Gabrius ; characters; Daghestan; Caspian region ) .

— Coiffait, 1974: 69 ( Gabrius ; characters; Daghestan; Transcaspian region ) .

- DISTRIBUTION: Russia, Azerbaijan, “Transcaspian region”.

musonoiensis Levasseur, 1967: 655 ( Gabrius ; Type locality: région de Kolwezi, Katanga, Musonoie).

- DISTRIBUTION: Congo.

multipunctatus Blatchley, 1910 , see: ovaliceps Fall, 1930 .

multipunctatus Shibata, 1991 , see: schillhammeri Shibata, 1996 .

nanellus Casey, 1915 , see: nigritulus Gravenhorst, 1802 .

nanus Gravenhorst, 1806 , see: splendidulus Gravenhorst, 1802 .

negus Tottenham, 1956b: 217 ( Gabrius ; Type locality: Abyssinia: Djem-Djem Forest ).

- DISTRIBUTION: Ethiopia.

neobisniformis Bernhauer, 1915d: 144 ( Philonthus ; subgenus Gabrius ; Type locality: Abessinien: Dire Daoua).

— Scheerpeltz, 1933: 1352 ( Philonthus ; catalog).

— Schillhammer, 1997: 79 ( Gabrius ).

- DISTRIBUTION: Ethiopia.

nepalensis Schillhammer, 1997: 11 ( Gabrius ; Type locality: Nepal: [ Patan District ], Kathmandu District, Phulcoki, 2600 m).

- DISTRIBUTION: Nepal.

nepos Smetana, 1984: 146 ( Gabrius ; Type locality: Japan: Gumma Pref.: Usui Bypass , 700 m). - DISTRIBUTION: Japan.

neptunus Schillhammer, 1997: 43 ( Gabrius ; Type locality: India: W. Bengal, Darjeeling Distr. Ghoom-Lopchu, 2000 m. Also cited from Nepal).

— Schillhammer, 2000 c: 132 ( Gabrius ; Nepal).

— Bernhauer and Schubert, 1914: 347 ( Philonthus ; catalog).

— Johansen, 1914: 418 ( Philonthus ; subgenus Gabrius ; characters; Denmark).

— Notman, 1919a: 95 ( Philonthus ; Ontario).

— Gridelli, 1920: 142 ( Gabrius ; characters; Italy).

— Šulc, 1922 ( Gabrius ; osmeterium; anatomy; histology; physiology).

— Lea, 1925: 220 ( Philonthus ; Australian introduction).

— Fall, 1926: 150 ( Philonthus ; Alaska, Yukon Territory).

— Cameron and Scott, 1928: 272 ( Philonthus ; Rodriguez Island).

— Portevin, 1929: 366 ( Philonthus ; characters; France).

— Gridelli, 1930: 74 ( Gabrius ; Libya).

— Bernhauer, 1931d: 588 ( Philonthus ; Abyssinia).

— Peyerimhoff, 1931: 32 ( Philonthus ; subgenus Gabrius ; notes; Algeria).

— Cameron, 1932: 147 ( Philonthus ; characters; Ceylon).

— Scheerpeltz, 1933: 1352 ( Philonthus ; catalog).

— Wüsthoff, 1934: 61 ( Philonthus ; illustration of aedeagus).

— Scheerpeltz, 1935a: 654 ( Philonthus ; Australia).

— Uyttenboogaart, 1937: 82 ( Philonthus ; notes; Canary Islands).

— Bernhauer, 1940c: 4 ( Philonthus ; Azores; Madeira).

— C. Koch, 1941: 64 ( Philonthus ; subgenus Gabrius ; Morocco).

— Méquignon, 1942a: 21 ( Gabrius ; Azores).

— Fagel, 1945: 40 ( Gabrius ; characters; Belgium).

— Jeannel and Jarrige, 1949: 376 ( Philonthus ; subgenus Gabrius ; cave species; France; Serbia; Romania).

— Coiffait, 1951: 113, 115 ( Gabrius ; characters in key; France).

— V. Hansen, 1952: 89 ( Philonthus ; subgenus Gabrius ; characters; Denmark).

— Smetana, 1952b: 120 ( Gabrius ; characters; Czechoslovakia).

— Smetana, 1953: 122 ( Gabrius ; Turkey).

— Coiffait, 1953a: 265 ( Gabrius ; [Note: Misidentified as latro Joy according to Schillhammer, 1997: 80]; characters; France).

— Coiffait, 1954: 165 ( Gabrius ; Canary Islands).

— Scheerpeltz, 1956a: 485 ( Gabrius ; Belgian Congo).

— Smetana, 1957: 67, 69 ( Gabrius ; characters).

— Smetana, 1958: 248 ( Gabrius ; characters; ecology; Czechoslovakia).

— Larsson and Gígja, 1959: 87 ( Philonthus ; biological notes; Iceland).

— Smetana, 1960c: 325 ( Gabrius ; characters; wide-spread species).

— Scheerpeltz, 1961d: 130 ( Gabrius ; Italy).

— Smetana, 1961a: 44 ( Gabrius ; Bulgaria).

— Smetana, 1962c: 236 ( Gabrius ; notes on specimens in Gravenhorst collection; lectotype designation).

— Smetana, 1963b: 38 ( Gabrius ; Madeira).

— Palm, 1963b: 105 ( Philonthus ; subgenus Gabrius ; characters; Sweden; Denmark; Norway; Finland).

— Lohse, 1964: 191 ( Gabrius ; characters; central Europe).

— Scheerpeltz, 1964a: 300 ( Gabrius ; Crete).

— Focarile, 1964: 56 ( Gabrius ; Italy).

— Moore, 1965: 100 ( Gabrius ).

— Scheerpeltz, 1965b: 403 ( Gabrius ; Sicily).

— Scheerpeltz, 1965c: 507 ( Gabrius ; Sicily).

— Horion, 1965: 194 ( Gabrius ; all Europe).

— Pototskaia, 1966: 154 ( Gabrius ; larval characters).

— Pototskaia, 1967: 57 ( Gabrius ; larval characters).

— Smetana, 1967b: 159 ( Gabrius ; Spain).

— Smetana, 1967i: 556 ( Gabrius ; Turkey) .

— Smetana, 1970: 59 ( Gabrius ; distributional summary; Azores).

— Fagel, 1972: 103 ( Gabrius ; notes; Sainte Helene).

— Runde, 1835: 11 ( Staphylinus ; characters; Germany).

— Lacordaire, 1835: 408 ( Staphylinus ; cited as atterimus; characters; France).

— Nordmann, 1837: 126 ( Gyrohypnus ; Europe).

— Westwood, 1838a: 16 ( Gabrius ; characters; type species).

— Zetterstedt, 1838: 65 ( Staphylinus ; characters; habitat; Lapland).

— Erichson, 1839a: 476 ( Philonthus ; characters; Germany).

— Stephens, 1839: 400 ( Gabrius ; characters; England).

— Erichson, 1840: 492 ( Philonthus ; characters; Europe; America septentrionalis). — Laporte, 1840: 179 ( Cafius ; characters; France).

— P. Lucas, 1846: 113 ( Philonthus ; notes; Algeria).

— Jacobsen, 1849: 703 ( Philonthus ; characters; Denmark).

— Hochhuth, 1849: 144 ( Philonthus ; characters).

— Redtenbacher, 1849: 706 ( Philonthus ; characters; Austria).

— LeConte, 1850: 220 ( Philonthus ; Lake Superior).

— Hardy, 1851: 31 ( Philonthus ; notes; England).

— Mäklin, 1853: 188 ( Philonthus ; characters; Kadjak Island).

— Wollaston, 1854: 584 ( Philonthus ; characters; notes; Madeira Islands).

— Küster, 1854: 16 ( Philonthus ; characters; Europe).

— G. Waterhouse, 1858: 25 ( Philonthus ; synonym of nigritulus ).

— Rosenhauer, 1856: 71 ( Philonthus ; notes; Algeciras; Granada).

— Fairmaire and Laboulbène, 1856: 528 ( Philonthus ; characters; France).

— Kraatz, 1857d: 616 ( Philonthus ; synonym of nigritulus ).

— Redtenbacher, 1857: 196 ( Philonthus ; characters; Austria).

— Peyron, 1858: 427 ( Philonthus ; Turkey).

— Thomson, 1860: 170 ( Gabrius ; characters; Scandinavia).

— Gemminger and Harold, 1868: 590 ( Philonthus ; synonym of nigritulus ).

— Redtenbacher, 1874: 215 ( Philonthus ; characters; Austria).

— Ganglbauer, 1895: 460 ( Philonthus ; synonym of nigritulus ).

— Bernhauer and Schubert, 1914: 347 ( Philonthus ; synonym of nigritulus ).

— Cameron, 1932: 147 ( Philonthus ; synonym of nigritulus ).

— Scheerpeltz, 1933: 1352 ( Philonthus ; synonym of nigritulus ).

— Smetana, 1958: 249 ( Gabrius ; synonym of nigritulus ).

— Smetana, 1960c: 325 ( Gabrius ; synonym of nigritulus ).

— Fagel, 1972: 103 ( Gabrius ; synonym of nigritulus ).

— Coiffait, 1974: 54 ( Gabrius ; synonym of nigritulus ).

— Smetana, 1995e: 619 ( Gabrius ; synonym of nigritulus ).

pumilus Mannerheim, 1830: 32 ( Cafius ; Type locality: Ostrogothia Sveciae). — Mannerheim, 1831: 446 ( Cafius ; Sweden).

— Erichson, 1840: 492 ( Philonthus ; synonym of aterrimus ).

— Hochhuth, 1849: 144 ( Philonthus ; synonym of aterrimus ).

— Küster, 1854: 16 ( Philonthus ; synonym of aterrimus ).

— Kraatz, 1857d: 616 ( Philonthus ; synonym of nigritulus ).

— Motschulsky, 1858: 661 ( Gabrius ; characters; Germany).

— Gemminger and Harold, 1868: 591 ( Philonthus ; variety of nigritulus ).

— Fauvel, 1874: 243 [= 1874a: 469] ( Philonthus ; synonym of nigritulus ).

— Ganglbauer, 1895: 460 ( Philonthus ; synonym of nigritulus ).

— Bernhauer and Schubert, 1914: 348 ( Philonthus ; synonym of nigritulus ).

— Cameron, 1932: 147 ( Philonthus ; synonym of nigritulus ).

— Smetana, 1958: 249 ( Gabrius ; synonym of nigritulus ).

— Smetana, 1960c: 325 ( Gabrius ; synonym of nigritulus ).

— Coiffait, 1974: 54 ( Gabrius ; synonym of nigritulus ).

— Smetana, 1995e: 619 ( Gabrius ; synonym of nigritulus ).

pallipes Stephens, 1833: 250 ( Gabrius ; Type locality: Norfolk; Suffolk).

— Stephens, 1839: 400 ( Gabrius ; characters; England).

pygmaeus Stephens, 1833: 250 ( Gabrius ; Type locality: Suffolk; Norfolk; Devonshire). — Stephens, 1839: 400 ( Gabrius ; characters; England).

— G. Waterhouse, 1858: 25 ( Philonthus ; synonym of nigritulus ).

— Gemminger and Harold, 1868: 591 ( Philonthus ; synonym of nigritulus ).

phaeopus Stephens, 1833: 250 ( Gabrius ; Type locality: London; Hants.: New Forest). — Stephens, 1839: 400 ( Gabrius ; characters; England).

— G. Waterhouse, 1858: 25 ( Philonthus ; synonym of nigritulus ).

— Gemminger and Harold, 1868: 591 ( Philonthus ; synonym of nigritulus ).

— Bernhauer and Schubert, 1914: 347 ( Philonthus ; synonym of nigritulus ).

— Cameron, 1932: 147 ( Philonthus ; synonym of nigritulus ).

— Smetana, 1958: 249 ( Gabrius ; synonym of nigritulus ).

— Smetana, 1960c: 325 ( Gabrius ; synonym of nigritulus ).

— Coiffait, 1974: 54 ( Gabrius ; synonym of nigritulus ).

— Smetana, 1995e: 619 ( Gabrius ; synonym of nigritulus ).

attenuatus Stephens, 1833: 251 ( Gabrius ; Type locality: Battersea-fields; Devonshire). — Stephens, 1839: 401 ( Gabrius ; characters; England).

— G. Waterhouse, 1858: 25 ( Philonthus ; synonym of nigritulus ).

— Gemminger and Harold, 1868: 590 ( Philonthus ; synonym of nigritulus ).

— Bernhauer and Schubert, 1914: 347 ( Philonthus ; synonym of nigritulus ).

— Cameron, 1932: 147 ( Philonthus ; synonym of nigritulus ).

— Smetana, 1958: 249 ( Gabrius ; synonym of nigritulus ).

— Smetana, 1960c: 325 ( Gabrius ; synonym of nigritulus ).

— Coiffait, 1974: 54 ( Gabrius ; synonym of nigritulus ).

— Smetana, 1995e: 619 ( Gabrius ; synonym of nigritulus ).

basalis Stephens, 1833: 251 ( Gabrius ; Type locality: London).

— Stephens, 1839: 400 ( Gabrius ; characters; England).

— G. Waterhouse, 1858: 25 ( Philonthus ; synonym of nigritulus ).

— Gemminger and Harold, 1868: 590 ( Philonthus ; synonym of nigritulus ).

— Bernhauer and Schubert, 1914: 347 ( Philonthus ; synonym of nigritulus ).

— Cameron, 1932: 147 ( Philonthus ; synonym of nigritulus ).

— Smetana, 1958: 249 ( Gabrius ; synonym of nigritulus ).

— Smetana, 1960c: 325 ( Gabrius ; synonym of nigritulus ).

— Coiffait, 1974: 54 ( Gabrius ; synonym of nigritulus ).

— Smetana, 1995e: 619 ( Gabrius ; synonym of nigritulus ).

pusillus Heer, 1839: 271 ( Philonthus ; Type locality: Bern).

— Kraatz, 1857d: 610 ( Philonthus ; characters; Germany).

— Fauvel, 1874: 243 [= 1874a: 469] ( Philonthus ; synonym of nigritulus ).

— Ganglbauer, 1895: 460 ( Philonthus ; synonym of nigritulus ).

— Bernhauer and Schubert, 1914: 348 ( Philonthus ; synonym of nigritulus ).

— Cameron, 1932: 147 ( Philonthus ; synonym of nigritulus ).

— Smetana, 1958: 249 ( Gabrius ; synonym of nigritulus ).

— Smetana, 1960c: 325 ( Gabrius ; synonym of nigritulus ).

— Coiffait, 1974: 54 ( Gabrius ; synonym of nigritulus ).

— Smetana, 1995e: 619 ( Gabrius ; synonym of nigritulus ).

pygmaeus Snellen, 1854: 70 ( Staphylinus ; [preoccupied]; Type locality: Nederland). — Gemminger and Harold, 1868: 591 ( Philonthus ; variety of nigritulus ).

— Fauvel, 1869: 488 ( Philonthus ; synonym of nigritulus ).

— Fauvel, 1874: 243 [= 1874a: 469] ( Philonthus ; synonym of nigritulus ).

— Ganglbauer, 1895: 460 ( Philonthus ; synonym of nigritulus ).

— Bernhauer and Schubert, 1914: 348 ( Philonthus ; synonym of nigritulus ).

— Cameron, 1932: 147 ( Philonthus ; synonym of nigritulus ).

— Smetana, 1958: 249 ( Gabrius ; synonym of nigritulus ).

— Smetana, 1960c: 325 ( Gabrius ; synonym of nigritulus ).