Sanaungulus marginalis, Yang & Zhao & Liu, 2024

Yang, Yuxia, Zhao, Wei & Liu, Haoyu, 2024, Taxonomic notes and new species of Burmomiles and Sanaungulus (Coleoptera, Cantharidae) from northern Myanmar during the late Mesozoic, Contributions to Zoology 2024 (2), pp. 153-177 : 159-160

Sanaungulus marginalis sp. nov.

(fig. 1)

Remarks. In the original description, it was Etymology. The specific epithets derived from described as follows “sex undefined, anten- Latin “ margin ” (margin), referring to its pronomere V … at apex with an extremely long notum with nearly straight lateral margins. antennal process…; VI … with at apex an anten- Material. Currently only one specimen nal process…; antennomere VII … with an is known. This holotype is deposited in the apical process …; VIII–IX … with an apical pro- Museum of Hebei University, Baoding, China cess…” (Bramanti & Fanti, 2022). According to (MHBU, No. BU-COL-CAN0013).

the shape of the antennae, it should be a male, Horizon and locality. The lowermost and actually the antennomeres VI–X each has Cenomanian (98.79 ± 0.62 Ma), mid-Creta- an antennal appendage near the base. ceous (Shi et al., 2012). Myanmar: Kachin State, Myitkyina District, Tanai Township, Hukawng Valley .

Sanaungulus perkovskyi Fanti & Müller, Differential diagnosis: The species looks

2022 similar to S. multiramus Yang, Liu & Zhao , Sanaungulus perkovskyi Fanti & Müller, 2022: 2022 and S. ruficollis Yang Liu & Zhao, 2022

365, fig. 19. in antennomeres IV –X each with an antennal appendage, triangular head and elliptical Remarks. In their original publication, eyes, but can be distinguished from the latter Fanti & Müller (2022) described that “male, two by the shape of pronotum, which is traantennomeres IV – VI extremely long, each pezium, obliquely narrowed anteriorly, with equipped with an antennal process at apex…; all margins nearly straight. In comparison, the antennomere VII … equipped with a long pronotum of S. multiramus and S. ruficollis is antennal process; antennomere VIII …with- suborbicular, anterior margin rounded, latout the antennal process…”. As evidenced eral margins Downloaded arcuate from and Brill.com slightly converging 06/21/2024 06:10:25 PM via Open Access. This is an open access article distributed under the terms of the CC BY 4.0 license. https://creativecommons.org/licenses/by/4.0 /

anteriorly, posterior margin rounded in S. multiramus , almost straight in S. ruficollis (Yang et al., 2022: figs. 7, 8, 14, 15).

Description. Male (fig. 1). Total length 3.80 mm (from anterior margin of clypeus to apices of elytra); width 1.10 mm. Body approximately 3.5 times longer than wide. Head and pronotum yellow brown, antennae, elytra and legs dark brown.

Head (fig. 1A) 0.67 mm long, 0.88 mm wide, triangular. Eyes large and prominent, elliptical, inter-ocular distance approximately 2.5 times greater than the eye diameter. Mandibles with a tooth near apical one-third part of inner edge. Terminal maxillary palpomere and terminal labial palpomere not well visible. Antennae (fig. 1A, B) pectinate, probably surpassing elytral mid-length if extending posteriorly, scape stout and distinctly longer than pedicel, pedicel as long as III and expanding apically, IV distinctly longer than III, IV–X parallel-sided and each with an antennal appendage inserted at a basal cavity on the ventral side, XI bilaterally compressed and widened; relative length of antennomeres as 10:2:3:16:16:16:16:17:15:14:16.

Pronotum (fig.1A, B) 0.61 mm long, 0.85 mm wide, nearly as wide as head, obliquely narrowed anteriorly, all margins nearly straight, anterior angles and posterior angles rounded. Scutellum triangular, nearly twice as wide as long, rounded at apical.

Elytra (fig. 1A, C) 2.80 mm long, 1.10 mm wide, at base slightly wider than pronotum, approximately 2.5 times as long as width at humeri, lateral margins parallel and simple, apices rounded, surface finely punctate, punctures deep and rounded in striate, which distributed along with the moderately indicated longitudinal costae on disc. Hind wing distinctively longer than elytra when unfolded.

Legs (fig. 1A) not well visible, at least left meta-tarsomere I twice longer than tarsomere II.

Abdomen not well visible.

Remarks. The yellow amber piece originally measured 15 × 13 × 5 mm. The legs and abdomen were not well visible in the amber.