Parachorius javanus ( Boucomont, 1914 ) Tarasov, 2017

Tarasov, Sergei, 2017, A cybertaxonomic revision of the new dung beetle tribe Parachoriini (Coleoptera: Scarabaeidae: Scarabaeinae) and its phylogenetic assessment using molecular and morphological data, Zootaxa 4329 (2), pp. 101-149 : 128-129

Parachorius javanus ( Boucomont, 1914) new combination

( Figs. 14 , 26C )

Cassolus javanus Boucomont, 1914: 251 Cassolus javanus ; Balthasar, 1963: 263

Type locality. Sokabumi [West Java].

Diagnosis. Along with P. schuelkei and P. pseudojavanus , this species can be separated from all other Parachorius the two distinctly produced clypeal teeth whose outer margin is not notched basally. Parachorius javanus can be distinguished from P. schuelkei and P. pseudojavanus by a combination of characters: (1) body coloration is reddish brown, and (2) parameres are notched basally and claw shaped in lateral view. External morphology and endophallic sclerite characters place P. javanus as most closely related to P. pseudojavanus .

Taxonomic notes. The species is known only from the holotype. All other records of P. javanus are likely to refer to the newly described and closely related P. pseudojavanus . Parachorius javanus and P. pseudojavanus can only be reliably separated by the shape of aedeagus; the females of the two species are undistinguishable. Due to this, many females from various localities in West Java remain unidentified.

Description. Body. Body reddish brown, length 6.7 mm. Elytra uniformly colored.

Head. Clypeus with two distinctly produced teeth, their outer margin not notched basally.

Pronotum. Pronotum rounded laterally. Frontal angles rounded. Disc covered with punctures separated by 1–4 puncture diameter(s). Pronotum in anterior-posterior direction notably shorter than elytra.

Elytra. Elytra covered with sparse punctures.

Wings. Wings normally developed.

Legs. Profemora with almost straight or sinuate anterior margin. Protibia with apical spur approximately reaching middle of protarsomere 3. First tooth notably modified in males, approximately twice as wide as second tooth; first and second teeth distinctly separated from each other; first tooth distinctly separated from denticles located between first and second teeth. Metafemoral posterior margin not serrate, simple, not modified. Metatibia slightly sinuate, inner margin not denticulate, straight, not produced apically.

Metasternum. Transverse basal ridge of metasternum long, at least 1/5 of mesocoxa length.

Aedeagus. Parameres symmetrical, notched basally, and claw shaped in lateral view.

Material examined. Holotype: 1 male, " Java occident Sukabumi 2000 1983 H. Fruhstorfer / EX MUSEO N . VAN DE POLL] / TYPUS / Boucomont det. 1914 Cassolus javanus n.sp. / MUSEUM PARIS Boucomont / J. Krikken 1969 compared with 5 PM ( Amsterdam : ZMUA) which is comp[letely]. Rufous from Bogor ", 6°55.1'S, 106°55.6'E ( MNHN) .

Figures

FIGURE 14. Morphology of Parachorius javanus. (A, E) aedeagus lateral view (male, holotype). (B) habitus (male, holotype). (C – D) protibia (male, holotype)

FIGURE 26. Distribution maps of Parachorius species. (A) Parachorius hookeri. (B) Parachorius humeralis. (C) Parachorius javanus. (D) Parachorius longipenis. (E) Parachorius maruyamai. (F) Parachorius newthayerae. Type locality — type locality of species (holotype or lectotype); Type. Collection data — localities of type specimens associated with species (paratypes or paralectotypes); Collection data — localities of other specimens examined in this study.