Pseudotorinia grasemanni, Harzhauser & Landau, 2023

Harzhauser, Mathias & Landau, Bernard, 2023, The Architectonicidae and Mathildidae (Gastropoda, Heterobranchia) of the Miocene Paratethys Sea-victims of the Miocene Climatic Transition, Zootaxa 5370 (1), pp. 1-74 : 39-40

Pseudotorinia grasemanni nov. sp.

Figs 4N , 19A–D

Architectonica (Nipteraxis) marthae Boettger — Csepreghy-Meznerics 1956: 386, pl. 1, figs 19–21 [non Pseudotorinia marthae ( Boettger, 1902) ].

Architectonica (Nipteraxis) martae [sic] (Boettger)—Kojumdgieva in Kojumdgieva & Strachimirov 1960: 93, pl. 29, figs 17a–d [non Pseudotorinia marthae ( Boettger, 1902) ].

Solarium marthae Boettger —Strausz 1962: 122, pl. 51, figs 5–7 [non Pseudotorinia marthae ( Boettger, 1902) ].

Architectonica (Nipteraxia [sic]) marthae Boettger, 1901 —Strausz 1966: 118, pl. 51, figs 5–7 [non Pseudotorinia marthae ( Boettger, 1902) ].

Solarium marthae Boettger — Csepreghy-Meznerics 1969b: 20, pl. 2, figs 19, 23, 26 [non Pseudotorinia marthae ( Boettger, 1902) ].

Type material. Holotype: NHMW 2023 /0267/0007, SL: 2.8 mm, MD: 6.1 mm, Steinebrunn ( Austria), Figs 19B 1 – B 3 . Paratypes: NHMW 2023 /0267/0008, SL: 2.7 mm, MD: 7.0 mm, Steinebrunn ( Austria), Figs 19A 1 –A 3 . NHMW 2023 /0267/0009, SL: 2.5 mm, MD: 6.0 mm, Steinebrunn ( Austria), Figs 19C 1 –C 3 . NHMW 2023 /0267/0010, SL: 2.9 mm, MD: 5.8 mm, Steinebrunn ( Austria), Figs 4N , 19D 1 –D 3 .

Type locality. Steinebrunn ( Austria), Vienna Basin .

Type stratum. Silty sand of the Baden Formation.

Age. Middle Miocene, early/middle Badenian (Langhian).

Etymology. In honor of Bernhard Grasemann, Geologist at the Department of Geology of the University of Vienna ( Austria).

Diagnosis. Small, depressed lentiform shell with low conical to nearly flat spire and angled periphery with rounded keel. Dorsal sculpture of finely beaded spiral cords with SSC bearing slightly more prominent beads. Base with wide umbilicus and succession of five delicate beaded spiral threads on outer basal field and three stronger beaded spiral cords on the inner basal field.

Description. Small, depressed lentiform shell with low conical to nearly flat spire. Protoconch slightly submerged below apex, small, of <1 visible whorl (diameter 0.6 mm). Teleoconch of ~3.5 whorls. First teleoconch whorl flat with slightly raised, beaded SSC and UPC with two finer beaded MCs. Later whorls with very prominent SSC bearing rounded beads, three weaker MCs and prominent UPC. LPC separated from UPC by narrow but slightly deeper groove; largely covered by subsequent whorl causing relatively deep suture. Beads densely spaced, axially elongated, arranged in prosocline lines. LPC forming blunt keel; below steeply inclined nearly flat to faintly concave passing via weak angulation into weakly convex base. Periphery finely beaded in basal view. No IPC developed. Five narrow, densely beaded spiral cords on outer basal field and three broader beaded cords on inner basal field (innermost corresponds to PUC). Growth lines forming weakly cancellate pattern in interspaces between spiral cords. UC delimited from basal field by distinct groove, prominent, slightly sunken into and overhanging umbilicus. Umbilicus very wide (~31–38 of MD). Umbilical wall bearing fine axial growth lines. Aperture small, circular, CG and PG very weak.

Discussion. Pseudotorinia grasemanni is reminiscent of P. marthae ( Boettger, 1902) with which it has been confused in the literature. They can generally be separated by the wider umbilicus of P. grasemanni (~31–38 vs. ~20–26 of MD). Moreover, the beads on the periphery are coarser in P. marthae . There are several species described from the European Neogene, which agree with Pseudotorinia grasemanni in the wide umbilicus. Of these, the Pliocene to extant Pseudotorinia architae (O.G. Costa, 1841), from the Mediterraean Sea, might be a closely related species, but is distinguished by its comparatively uniform sculpture on the dorsal side and the rounded periphery, which is in a more adapical position ( Bieler 1993: fig. 225; Chirli 2013; pl. 6, figs 3–9). Pseudotorinia ivolasi ( Ivolas & Peyrot, 1900) , from the Langhian of the Loire Basin ( France), differs in its nearly flat spire and rounded periphery ( Glibert 1949 pl. 6, figs 15a-b, pl. 7, fig. 8). Pseudotorinia planulatum ( Grateloup, 1832) , from the Burdigalian of the Aquitaine Basin ( France), differs in its axial sculpture on the basal field and the reticulate sculpture on the dorsum ( Cossmann & Peyrot 1919: pl. 15, figs 71–73; https://science.mnhn.fr/institution/mnhn/collection/f/ item/j05829?listIndex=68&listCount=169). Pseudotorinia subplicata (d’Orbigny, 1852) , from the Burdigalian of the Aquitaine Basin ( France), differs in its rounded periphery ( Cossmann & Peyrot 1919: pl. 15, figs 59–61). The Pliocene Mediterranean Pseudotorinia obtusa ( Bronn, 1831) differs in its narrower umbilicus, more rounded periphery, and more elevated spire ( Ferrero Mortara et al. 1984 pl. 15, figs 4a–c).

Distribution. Known so far only from the middle Badenian (late Langhian) of the Central Paratethys Sea.

Central Paratethys Sea. Middle Miocene (Badenian): Vienna Basin: Baden, Steinebrunn ( Austria) (hoc opus); Pannonian Basin: Szob ( Hungary) (Strausz 1966); B̧kk Mountains: Sámsonháza ( Hungary) ( Csepreghy-Meznerics 1969b); Dacian Basin: Opanec ( Bulgaria) (Kojumdgieva in Kojumdgieva & Strachimirov 1960).

Figures

FIGURE 1. Descriptive terms used in descriptions of Architectonicidae (modified from Bieler 1993).

FIGURE 3. Descriptive terms used in descriptions of Mathildidae.

FIGURE 4 Protoconchs of Paratethyan Architectonicidae. A. Ammotectonica gregorovae nov. sp., paratype, NHMW 1862/0029/0078c. B. Simplexollata anticollata nov. sp., paratype, NHMW 1854/0035/0331, Lăpugiu de Sus (Romania). C. Simplexollata carocollata (Lamarck, 1822), NHMW 1850/0018/0115, Léognan (France), Burdigalian. D. Simplexollata simplex (Bronn, 1831), NHMW 2013/0078/0109, Baden (Austria).E. Nipteraxis deformatusnov. sp., paratype, NHMW 2010/0004/0456a, Bad Vöslau (Austria). F. Nipteraxis exmoniliferus (Sacco, 1892), NHMW 1907/0323/1959, M̂llersdorf (Austria). G. Solatectonica kostejana (Boettger, 1907), NHMW 2012/0229/0049b, Traiskirchen (Austria). H. Nipteraxis monilifer (Bronn, 1831), NHMW 1846/0037/0427, M̂llersdorf (Austria). I. Solatisonax tavianii nov. sp., NHMW2016/0266/0002, SL: 8.0 mm, MD: 15.0 mm, Lăpugiu des Sus (Romania). J. Solatisonax transversa nov. sp., holotype, NHMW 2016/0177/0341, Lăpugiu de Sus (Roma- nia). K. Granosolarium semilaevis (Sacco, 1892), NHMW 1846/0037/0428d, M̂llersdorf (Austria). L. Heliacus globosus nov. sp., holotype NHMW 1907/0328/1959a, M̂llersdorf (Austria). M. Heliacus miser (Dujardin, 1837), NHMW 2023/0267/0001, Steinebrunn (Austria). N. Pseudotorinia grasemanni nov. sp., paratype, NHMW 2023/0267/0010, Steinebrunn (Austria). O. Pseudotorinia marthae (Boettger, 1902), NHMW 2012/0229/0049, Traiskirchen (Austria). All figures same scale.

FIGURE 19 A1–A3. Pseudotorinia grasemanni nov. sp., paratype, NHMW 2023/0267/0008, Steinebrunn (Austria). B1–B3. Pseudotorinia grasemanni nov. sp., holotype, NHMW 2023/0267/0001, Steinebrunn (Austria). C1–C3. Pseudotorinia grasemanni nov. sp., paratype, NHMW 2023/0267/0009, Steinebrunn (Austria). D1–D3. Pseudotorinia grasemanni nov. sp., paratype, NHMW 2023/0267/0010, Steinebrunn (Austria).