Nipteraxis exmoniliferus (Sacco, 1892)

Harzhauser, Mathias & Landau, Bernard, 2023, The Architectonicidae and Mathildidae (Gastropoda, Heterobranchia) of the Miocene Paratethys Sea-victims of the Miocene Climatic Transition, Zootaxa 5370 (1), pp. 1-74 : 34-35

publication ID	https://doi.org/ 10.11646/zootaxa.5370.1.1
publication LSID	lsid:zoobank.org:pub:48903495-7C6C-46E4-9B1B-D6A2F2781873
DOI	https://doi.org/10.5281/zenodo.10580842
persistent identifier	https://treatment.plazi.org/id/03BEE17B-FFE0-C046-FF0E-FE6078B8304B
treatment provided by	Plazi
scientific name	Nipteraxis exmoniliferus (Sacco, 1892)
status

Treatment

Nipteraxis exmoniliferus (Sacco, 1892) View in CoL

Figs 4F View FIGURE 4 , 17A–D View FIGURE 17

Solarium moniliferum Bronn View in CoL — Ĥrnes 1848: 22 [non Niperaxis moniliferus ( Bronn, 1831) ].

Solarium moniliferum Bronn View in CoL —Ĥrnes 1856: 466, pl. 46, figs 5a–c [non Niperaxis moniliferus ( Bronn, 1831) ].

* [ Solarium (Solarium) carocollatum] var. exmonilifera Sacc. View in CoL —Sacco 1892: 43.

Solarium (Architectonica) moniliferum Bronn View in CoL —Boettger 1907: 135 [non Niperaxis moniliferus ( Bronn, 1831) ].

Solarium millegranum Lam. View in CoL (?)— Friedberg 1923: 414, pl. 25, fig. 6 [non Granosolarium millegranum ( Lamarck, 1822) View in CoL ].

Architectonica (Solariaxis) millegranum Lamk. — Korobkov 1955: plate captions, pl. 13, figs 19a–c. [non Granosolarium millegranum ( Lamarck, 1822) View in CoL ].

? Architectonica monilifera (Bronn) — Csepreghy-Meznerics 1956: 386, pl. 1, figs 29–30.

Architectonica (A [rchitectonica].) monilifera (Bronn) View in CoL —Sieber 1958: 139 [non Niperaxis moniliferus ( Bronn, 1831) ].

? Solarium moniliferum Bronn, 1831 View in CoL —Strausz 1966: 118, pl. 51, figs 8–10.

Solarium moniliferum Bronn View in CoL — Csepreghy-Meznerics 1969b: 20, pl. 2, figs 12, 17, 21, 27 [non Niperaxis moniliferus ( Bronn, 1831)].

Architectonica (Solariaxis) kostejanum (Boettger, 1906) — Atanacković 1985: 98, pl. 23, figs 14–17 [non Solatisonax kostejana (Boettger, 1907)].

exmonilifera (Solarium) Sacco, 1892 — Bieler & Petit 2005: 37.

non Architectonica (Architectonica) monilifera ( Bronn, 1831) — Atanacković 1985: 96, pl. 23, figs 1–3 [= Heliacus View in CoL ].

Type material. Holotype: NHMW 1851 View Materials /0013/0085, SL: 5.8 mm, MD: 9.2 mm, Steinebrunn ( Austria), illustrated by Ĥrnes (1856: pl. 46, fig. 5), Figs 17A View FIGURE 17 1 –A View FIGURE 1 2 View FIGURE 2 .

Illustrated material. NHMW 2007 View Materials z0078/0016a, SL: 6.3 mm, MD: 11.5 mm, Baden ( Austria), Figs 17B View FIGURE 17 1 –B View FIGURE 1 3 View FIGURE 3 . NHMW 2007 View Materials z0078/0016b, SL: 7.5 mm, MD: 12.5 mm, Baden ( Austria), Figs 17C View FIGURE 17 1 –C View FIGURE 1 3 View FIGURE 3 . NHMW 1864 View Materials /0001/0589, SL: 6.1 mm, MD: 9.3 mm, Steinebrunn ( Austria), Figs 17D View FIGURE 17 1 –D View FIGURE 1 3 View FIGURE 3 . NHMW 1907 View Materials /0323/1959, M̂llersdorf ( Austria), Fig. 4F View FIGURE 4 .

Additional material. 4 spec., NHMW 2012 View Materials /0213/0032, Pfaffstätten ( Austria) ; 1 spec., NHMW 2023 View Materials /0265/0001, M̂llersdorf ( Austria) ; 3 spec., NHMW 2023 View Materials /0268/0001, Baden ( Austria) ; 3 spec., NHMW 2023 View Materials /0300/0082, Baden-Sooss ( Austria) ; 2 spec., NHMW 2013 View Materials /0300/0081, Baden-Sooss ( Austria) ; 3 spec., NHMW 2022 View Materials /0205/0009, Vienna / Grinzing ( Austria) ; 3 spec., NHMW 1858 View Materials /0015/0190, Vienna / Pötzleinsdorf ( Austria) .

Description. Small to medium-sized solid shell with relatively high, cyrtoconoid spire, periphery with blunt keel. Protoconch heterostrophic, medium-sized of 1.2 visible whorls (diameter 1.05 mm). Transition to teleoconch marked by protoconch lip. Teleoconch of about 3.75 whorls. First teleoconch whorl flat with two prominent peripheral cords and dense axial ribs. Later whorls weakly convex. Sculpture of moderately prominent beaded SSC, three slightly weaker, beaded MCs and prominent, beaded UPC and LPC. Beads and spiral cords densely spaced. Periphery formed by prominent, convex LPC. Base faintly concave below keel; convex below. Densely beaded lower peripheral spiral cord and weaker IPC on base with weak, beaded spiral cord intercalated between UPC and LPC. Six spiral cords on basal field widening towards umbilicus. Innermost two cords (including PUC) forming relatively broad bands with axially elongated beads. UC coarse, prominent, overhanging umbilicus, delimited from PUC by narrow, moderately deep groove. Umbilicus narrowly funnel-shaped, wide (~26% of total diameter). Umbilical wall with distinct, prosocline growth lines and narrow central spiral cord. Aperture subcircular with narrow CG. PG absent.

Discussion. Ĥrnes (1856) and subsequent authors identified this species as Nipteraxis monilifer ( Bronn, 1831) , known from the Middle Miocene to Pliocene (Proto-)Mediterranean Sea, but also occurring in the Central Paratethys Sea (hoc opus). However, N. monilifer differs quite distinctly from N. exmoniliferus by its finely granulose spiral sculpture on the dorsal side, the more prominent keel, the well-defined LPC on the base, the relatively equal width of the spiral cords on the base, the deeper groove delimiting the UC and slightly wider umbilicus (see Ferrero Mortara et al. 1984; pl. 14, figs 5a–c). Nipteraxis exmoniliferus is reminiscent of Solatectonica kostejana (Boettger, 1907) , with which it was mixed in the NHMW collections. Both species are distinguished by the broad spiral cords of axial ribs on the basal field of S. kostejana and the more elevated spire of N. exmoniliferus . The Pliocene to Recent Mediterraean Solatisonax contexta (Seguenza, 1876) is a morphologically similar. It differs in the carinate keel, higher base, the weaker but broader spiral cords on the basal field, which lack prominent beads, and the narrower, more cylindric umbilicus (e.g., Seguenza 1903: 457, pl. 17, figs 3–5; Melone & Taviani 1982: 531, figs 1–3; Melone & Taviani 1985: 178, figs 48–52; Cauli & Bogi 2000: 12, pl. 2, figs 1–2; Bogi et al. 2002: 33, figs 12–13; Forli et al. 2002: 143, figs 21–23).

Ĥlzl (1973) listed Architectonica monilifera (Bronn) from the Late Oligocene of Bavaria. We doubt that this occurrence can be reliably identified and exclude it from the chresonymy herein.

Paleoenvironment. Shallow marine inner neritic; the occurrence at Steinebrunn ( Austria) indicates seagrass environments (own data M.H.). Occurrence in the clay of the Baden Fm. (e.g., Baden) may also point to deeper water habitats.

Distribution. Only known from the Badenian in the Central Paratethys Sea, where it occurred from the early to the late Badenian.

Central Paratethys Sea. Badenian (Middle Miocene): Polish-Ukrainian Carpathian Foredeep: Zboriv ( Ukraine) ( Friedberg 1923); Vienna Basin: Baden, Baden-Sooss, Möllersdorf, Pfaffstätten, Steinebrunn, Vienna / Grinzing, Vienna /Pötzleinsdorf ( Austria) (hoc opus, Sieber 1953); Pannonian Basin: Szob ( Hungary) ( Csepreghy-Meznerics 1956), Hrvaćani ( Bosnia and Herzegovina) ( Atanacković 1985); B̧kk Mountains: Borsodbóta ( Hungary) ( Csepreghy-Meznerics 1969b); Făget Basin: Coşteiu de Sus ( Romania) (Boettger 1907).