Pachydema

Micó, Estefanía & Galante, Eduardo, 2009, A new species of Pachydema Laporte (Coleoptera: Scarabaeoidea: Scarabaeidae: Melolonthinae: Pachydemini) from the Canary Islands, Zootaxa 2284, pp. 41-47 : 45-46

Key to the males of Pachydema species of the Canary Islands

All species of Canarian Pachydema are included in the key with the exception of P. integra (Wollaston) , which was described using a single female. The species distribution (modified from Lacroix 2000) is shown in brackets. Locality records in bold have not been verified by us.

1. Prementum convex ( Fig. 11 a). Maxillary teeth reduced, with setae or spine-like setae at apex ( Fig. 12 a). .............. 2

- Prementum showing a deep concave area at apical third ( Figs. 11 b–c). Maxillary teeth well developed ( Fig. 12 b).. ..................................................................................................................................................................................... 6

2. Antennal club more than twice as long the preceding 4 antennomeres combined ( Fig. 13 a). Genae protruding beyond canthus ( Fig. 14 a). ......................................................................................................................................... 3

- Antennal club less than 1.5 times the preceding 4 antennomeres combined ( Fig. 13 b). Genae not protruding beyond canthus (14b). [La Gomera]. ..................................................................................................... P. oromii López-Colón

3. Genae sharply angled and protruding beyond canthus ( Fig. 14 a). Parameres as in Fig. 16 a. [Tenerife, La Gomera, Fuerteventura]. ............................................................................................................................... P. obscura (Brullé)

- Genae rounded and slightly protruding beyond canthus. ............................................................................................ 3

4. Second protarsomere not widened (1.8 times longer than wide) Parameres as in Fig. 16 b. [Tenerife]. .......................

................................................................................................................................. P. tinerfensis Galante & Stebnicka - Second protarsomere slightly widened (1.3 times longer than wide). ......................................................................... 5

5. Elytra conspicuously convex and slightly punctate. Anterior border of clypeus emarginate. Parameres as in Fig. 16 c. [Tenerife]. ................................................................................................................................. P. fortunatorum Baraud

- Elytra less convex. Anterior border of clypeus very slightly emarginate. Parameres as in Fig. 16 d. [Tenerife]. ......... ............................................................................................................................................................ P. jeannei Baraud

6. Scutellum with punctures at disc. Metatibia internal border not dilated at middle ( Fig. 17 a). Parameres forcep shaped (narrow and sinuate, as in Figs. 16 e–h). ...................................................................................................................... 7

- Scutellum without punctures at disc. Metatibia internal border dilated at middle. Parameres not sinuate or arrow shaped (as in Figs. 16 i–n). ......................................................................................................................................... 10

7. Eyes large and laterally exposed. Front between eyes nearly twice as wide as eye diameter ( Fig. 14 c). [Tenerife, La Gomera]. ....................................................................................................................................... P. castanea (Brullé)

- Eyes small and not laterally exposed. Front between eyes more than three times as wide as eye diameter. .............. 8

8. Apical maxillary palpomere narrowly oval ( Fig. 15 a). Parameres as in Fig. 16 f. [Gran Canaria, Tenerife, La Palma, Fuerteventura, La Gomera]. .................................................................................................. P. fuscipennis (Brullé)

- Apical maxillary palpomere largely oval as in Fig. 15 b. ............................................................................................. 9

9. Frons and pronotum densely punctate. Parameres as in Fig. 16 g. [El Hierro, La Palma]. ... P. obscurella (Wollaston)

- Frons and pronotum less densely punctate. Parameres as in Fig. 16 h. [La Gomera]. ........... P. gomerae López-Colón

10. Eyes large and laterally exposed. Front between eyes nearly twice as wide as eye diameter. .................................. 11

- Eyes small and not laterally exposed. Front between eyes more than three times as wide as eye diameter. .......... 12

11. Apical maxillary palpomere large (ratio palpomere length/habitus length: 0.03) and widened (length 2.2 times the width). Third and fourth antennomeres short and wide (1.7 times longer than wide) ( Fig. 2 ). Parameres slightly arrow shaped ( Fig. 10 ). [Gran Canaria]. .............................................................................. P. megalops Micó sp. nov.

- Apical maxillary palpomere small (ratio palpomere length/habitus length: 0.01) and narrow (length 3.5 times the width). Third and fourth antennomeres long and narrow (3 times longer than wide) ( Fig. 13 c). Parameres strongly arrow shaped ( Fig. 16 i). [Gran Canaria]. ..................................................................................... P. lesnei Peyerimhoff

12. Second protarsomere strongly widened (almost as long as wide). Prementum widely notched and with the concave area very reduced ( Fig. 11 c). Parameres as in Fig. 16 j. [Gran Canaria]. ............................................. P. keithi Lacroix

- Second protarsomere moderately widened (always longer than wide). Prementum shortly notched, with a wide concave area in the apical third as in Fig. 12 b. ................................................................................................................ 13

13. Apical maxillary palpomere narrowly oval ( Fig. 15 e). Parameres as in Fig. 16 k. [Lanzarote, Fuerteventura]. ........... ........................................................................................................................................................... P. m e n i e r i Baraud

- Apical maxillary palpomere largely oval (Figs. f–h). ................................................................................................ 14

14. Parameres slightly arrow shaped ( Fig.16 l). Scutellum bordered by more or less deeply impressed groove. [El Hierro, Gran Canaria, Tenerife, La Gomera]. ........................................................................... P. bipartita (Brullé)

- Parameres elongated. Scutellum without marginal groove or only present minimally at the base. .......................... 15

15. Pronotum with fine, sparse punctures. Parameres as in Fig. 16 n. [Gran Canaria, Lanzarote]. ..... P. navatteae Baraud

- Pronotal punctures moderate to coarse, densely distributed. Parameres as in Fig. 16 m. [Fuerteventura]. ................... ............................................................................................................................................... P. wollastoni Peyerimhoff

Figures

FIGURES 11 – 15. 11 — Mentum: a) P. ob s cu ra, b) P. ob s cu rell a, c) P. keithi. 12 — Galea and lacinia of maxilla: a) P. obscura, b) P. obscurella. 13 — Antenna: a) P. o b s c u r a, b) P. oromi, c) P. l e s n e i. 14 — Head: a) P. o b s c u r a, b) P. oromi, c) P. castanea. 15 — Apical maxillary palpomere: a) P. fuscipennis, b) P. o b s c u re l l a, c) P. l e s n e i, d) P. keithi, e) P. m e n i e r i, f) P. wollastoni, g) P. bipartita, h) P. navatteae. Scale bar in mm.

FIGURES 16 – 17. 16 — Parameres, dorsal view: a) P. obscura, b) P. tinerfensis, c) P. fortunatorum, d) P. jeannei, e) P. castanea, f) P. fuscipennis, g) P. obscurella, h) P. g o m er a e, i) P. l e s n e i, j) P. keithi, k) P. m e n i e r i, l) P. bipartita, m) P. wollastoni, n) P. navatteae. 17 — Metatibia. a) P. obscurella, b) P. ke i t h i. Scale bar in mm.

FIGURES 1 – 10. Pachydema megalops Micó sp. nov. 1 — Head. 2 — Antenna. 3 — Galea and lacinia of maxilla. 4 — Apical maxillary palpomere. 5 — Mentum. 6 — Pronotum. 7 — Scutellum. 8 — Metatibia. 9 — Protarsus. 10 — Parameres, dorsal view. Scale bar in mm.