Chiromachetes Pocock, 1899

Sulakhe, Shauri, Deshpande, Shubhankar, Dandekar, Nikhil, Ketkar, Makarand, Gowande, Gaurang, Padhye, Anand & Bastawade, Deshabhushan, 2020, Two new species of Chiromachetes (Scorpiones: Hormuridae) from the northern Western Ghats, India, Euscorpius 320, pp. 1-27 : 4

Genus Chiromachetes Pocock, 1899

( Figures 1–83 , Tables 1–7 )

http: //zoobank. org/urn: lsid: zoobank. org: act: D1B455F7– E75B–4DF5–BAC7–31B0341D1784

TYPE SPECIES. Chiromachetes fergusoni Pocock, 1899 .

Figures

Figures 1–6. Chiromachetes parakrami sp. n., male, holotype, dorsal (1) and ventral (2) views, sternopectinal area (3), metasomal segment V and telson in lateral view (4), chelicera in dorsal view (5) and tarsi IV (6).

Figures 7–9. Chiromachetes parakrami sp. n., male, holotype, carapace (7), anterior margin of carapace (8) and left lateral eyes in dorsal view (9).

Figures 10–13. Chiromachetes parakrami sp. n., male, holotype in dorsal (10) and ventral (11) views, carapace (12) and dentition on movable finger of chela (13) under UV fluorescence.

Figures 14–18. Chiromachetes parakrami sp. n., female, paratype, BNHS SC 181, in dorsal (14) ventral (15) views, sternopectinal area (16), metasomal segment V and telson in lateral view (17) and carapace (18).

Figures 19–27: Chiromachetes parakrami sp. n. Figures 19–25. Male, holotype, pedipalp chela dorsal (19), external (20) and ventral (21) views, patella dorsal (22), ventral (23) and external (24) views and femur dorsal (25) view. Trichobothrial pattern indicated by yellow circles. Figures 26–27. Male, paratype, BNHS SC 180, right hemispermatophore in internal (26) and external (27) views.Abbreviations: p, pedicel; t, trunk; c, capsule; h, hook; dl, distal lamina.

Figures 28–30. Type locality of Chiromachetes parakrami sp. n., view of the dense semi–evergreen forest and mountains of Amba Ghat (28), view of the Amba Ghat road with adjacent boulders (29) and columnar basaltic bounder at the type locality from where the type series is collected (30).

Figure 31–36. Chiromachetes ramdasswamii sp. n., male, holotype, dorsal (31) and ventral (32) views, sternopectinal area (33), metasomal segment V and telson in lateral view (34), chelicera in dorsal view (55) and tarsi IV (36).

Figures 37–39. Chiromachetes ramdasswamii sp. n., male, holotype, carapace (37), anterior margin of carapace (38) and right lateral eyes in dorsal view (39).

Figures 40–43. Chiromachetes ramdasswamii sp. n., male, holotype in dorsal (40) and ventral (41) views, carapace (42) and dentition on movable finger of chela (43) under UV fluorescence.

Figures 44–48. Chiromachetes ramdasswamii sp. n., female, paratype, BNHS SC 184, in dorsal (44) ventral (45) views, sternopectinal area (46), metasomal segment V and telson in lateral view (47) and carapace (48).

Figures 49–57: Chiromachetes ramdasswamii sp. n. Figures 49–55. Male, holotype, pedipalp chela dorsal (49), external (50) and ventral (51) views, patella dorsal (52), ventral (53) and external (54) views and femur dorsal (55) view. Trichpobothrial pattern indicated by yellow circles. Figures 56–57. Male, paratype, INHER 254, right hemispermatophore in internal (56) and external (57) views. Abbreviations: p, pedicel; t, trunk; c, capsule; h, hook; dl, distal lamina.

Figures 58–60. Type locality of Chiromachetes ramdasswamii sp. n., view of riparian forest with waterfall at the type locality (58), view of Kawla fort through which the Varandha Ghat road is carved (59) and basaltic bounder adjacent to the Varandha Ghat road from where the type series is collected (60).

Figures: 61–63. Figure 61. Distribution of Chiromachetes in India.Figures 62–63. Chiromachetes sahyadriensis, male, INHER 172, carapace under UV light (62) and metasomal segment V and telson in lateral view (63).

Figures 64–69. Chiromachetes fergusoni, male topotype, dorsal (64) and ventral (65) views, sternopectinal area (66), metasoma V and telson lateral view (67), chelicera dorsal (68) and tarsi IV (69).

Figures 70–73. Chiromachetes fergusoni, male topotype, dorsal (70) and ventral (71) views under UV lights, carapace under white (72) and UV (73) light.

Figures 74–80. Chiromachetes fergusoni, male topotype, pedipalp chela dorsal (74), external (75) and ventral (76) views, patella dorsal (77), ventral (78) and external (79) views and femur dorsal (80) view. Trichobothrial pattern indicated by yellow circles.

Figures 81–82. Figure 81. Discriminant function analyses projection on first two factor planes explaining 96.85% of variation among the five species.Abbreviations:Circlecells (males), Squarecells (females).Figure 82. Maximum Likelihood phylogenetic tree (ML) for Chiromachetes from northern Western Ghats of India. Values along the nodes are bootstraps for 1000 iterations.

Figure 83. Bayesian phylogenetic tree for Chiromachetes from northern Western Ghats of India. Values along the nodes are Bayesian posterior probabilities for Bayesian Inference and Bayesian Poisson Tree Process (bPTP), respectively.