Cylindrinotina Español, 1956

Nabozhenko, Maxim & Ando, Kiyoshi, 2018, Subtribal, Generic And Subgeneric Composition Of Darkling Beetles Of The Tribe Helopini (Coleoptera: Tenebrionidae) In The Eastern Palaearctic Region, Acta Zoologica Academiae Scientiarum Hungaricae 64 (4), pp. 277-327 : 292-293

Subtribe Cylindrinotina Español, 1956

( Figs 7–26 )

General differential morphology. Adult. Head with ( Figs 14A , 18A ) or rarely without posterior grooves ( Figs 20A , 23A ) or with deep wrinkle ( Figs 8B , 16A ) beginning at lower aspect of the eyes. Labrofrontoclypeal membrane always exposed. Abdominal ventrite 5 not beaded along outer margin in most of all taxa excluding Microdocnemis Nabozhenko et Keskin, 2010 (but often with finely beaded posterior end).

Male genitalia. Apical piece flattened laterally at apex ( Figs 7B,F , 17B,G,L), with punctation, each puncture with very small seta directed toward the apex (‘nalassoid’ type of genitalia) or apical piece flattened dorso-ventrally at apex ( Figs 7G ), neither punctured nor setiferous (‘cylindrinotoid’ type of genitalia). Alae of apical piece moderately long, merged, concealed only half of median lobe, not reaching base of basal piece ( Figs 7A,E,I,O , 17A,F,K). Basal piece separated from alae by connecting membrane only in apical half, with connecting membrane between margins in basal half. Median lobe acute apically, only with one apex, with weakly sclerotized baculi, which are not connected at base. Female genital tubes. Spermatheca composed of single long tube and short accessory gland ( Figs 7M , 17P). Only the subgenus Caucasohelops Nabozhenko, 2006 of the genus Eustenomacidius Nabozhenko, 2006 has multibranched spermatheca (NABOZHENKO 2006). Valve between spermathecal and accessory gland ducts is absent.

Larva. Abdominal segment IX with small cylindrical or conical protuberances at base of urogomphi. Abdominal tergit VIII without protuberances.

Figures

Fig. 7. The genus Nalassus, genital structure: N. (s. str.) pilushenmuus (A–D), N. (s. str.) zoltani (E–H), N. (Nipponalassus) andoi (I–M), N. (Nipponalassus) lewisi (O–R): A, E, I, O = aedeagus, ventral view, B, F, J, P = aedeagus, lateral view, C, G, K, Q = male inner sternite VIII, D, H, L, R = gastral spicula, M = female genital tubes (ag – accessory gland of spermatheca,

Fig. 8. Nalassus (s. str.) brunneus brunneus, details of structure: A = head and prothorax, ventral view, B = head, ventral view (arrow shows weakly developed posterior grooves beginning at lower aspect of eyes), C = prosternum, D = prosternal process, procoxae and

Fig. 9. Nalassus brunneus konoi, holotype, female: A = habitus, B = labels, C = protibia, D = mesotibia

Fig. 10. Nalassus (s. str.) akitai, holotype and paratype: A = male, holotype, habitus, B = fe-

Fig. 11. Nalassus (s. str.) kawabatai, holotype and paratype. A = male, holotype, habitus, B = aedeagus, dorsal view, C = female paratype, habitus, D = male protibia, E = male mesotibia, F = male metatibia, G = male profemur, inner side, H = male prothorax, ventral view, I =

Fig. 12. Nalassus (s. str.) from Taiwan, habitus. A = N. formosanus, female, B = N. zoltani, male, C = N. pilushenmuus, male

Fig. 13. Nalassus (Nipponalassus) andoi, holotype and paratype. A = male, holotype, habitus, B = female paratype, habitus, C = male fore leg, dorsal margin, D = male profemur, inner side, E = aedeagus, dorsal view, F = prothorax, ventral view, G = abdominal ventrite 5, H = labels of the holotype

Fig. 14. Nalassus (Nipponalassus) andoi, details of structure. A = Head, ventral view, B = prothorax and base of fore legs, ventral side, C = male meso- and metaventrite, middle legs

Fig. 15. Nalassus (Nipponalassus) lewisi, holotype and paratype.A = male, holotype, habitus, B = female paratype, habitus, C = labels of the holotype, D = aedeagus, dorsal view

Fig. 16. Nalassus (Nipponalassus) lewisi (A, C–F) and N. (Nipponalassus) andoi (B), details of structure: A = head, ventral view, B = male abdominal ventrite 5, C = male meso- and metaventrite, middle legs basally, D = male abdominal ventrites, E = male profemur, inner side, F = male profemur dorsally

Fig. 18. Asialassus cordicollis, details of structure: A = head, ventral view, B = male prosternum, C = male prosternal process, D = female prothorax and fore legs, ventral view, E = female prosternal process, ventral view, F = female prothorax, posterior transverse view (lateral arrows show elevate margin of procoxae, middle arrow shows depressed prosternal process between procoxae), G = male protibia, H = male mesotibia

Fig. 19. Asialassus amamiensis, holotype, female: A = habitus, B = protibia, C = prothorax, ventral view, D = labels of the holotype

Fig. 20. Asialassus amamiensis, male, details of structure: A = head, B = prosternum, C = prosternal process, ventral view, D = prosternal process, lateral view

Fig. 21. Asialassus kimurai, holotype and paratype: A = holotype, male, habitus, B = paratype, female, habitus, C = male protibia, dorsally, D = male protibia, ventrally, E = male mesotibia, F = male prothorax ventro-laterally, G = male prothorax, ventrally, H = aedea-

Fig. 22. Asialassus kimurai, holotype and paratype of the new junior synonym Tarpela tokunoshimana: A = holotype, male, habitus, B = paratype, female, habitus, C = male prothorax, ventrally, D = male prothorax, laterally, E = male protibia, dorsally, F = male protibia, ventrally, G = male mesotibia, dorsally, H = male mesotibia, ventrally, I = labels of the holo-

Fig. 23. Asialassus kimurai, details of structure, male: A = head, ventral view, B = prothorax, ventral view, C = prosternal process, D = protibia, ventrally

Fig. 24. Asialassus tsushimanus, male, holotype: A = habitus, B = protibia, laterally, C = protibia, ventrally, D = mesotibia, E = aedeagus, dorsal view, F = prothorax, ventral view, G =

Fig. 25. Asialassus todai, male, holotype: A = habitus, B = prothorax, ventral view, C = prothorax, lateral view, D = protibia, dorsal view, E = protibia, ventral view, F = mesotibia, G =

Fig. 26. Asialassus adachii, male, holotype: A = habitus, B = protibia dorsally, C = protibia ventrally, D = mesotibia, E = prothorax, ventral view, F = prosternal process, lateral view, G = aedeagus, H = labels of the holotype