Symplecthyma Rivalier, 1964

Anichtchenko, Alexander & Wiesner, Jürgen, 2023, Philippine species of the genus Prothyma Hope, 1838 (Coleoptera: Cicindelidae). Part 2. Subgenus Symplecthyma Rivalier, 1964, Zootaxa 5357 (1), pp. 71-99 : 73

Key to the species of the subgenus Symplecthyma Rivalier, 1964

1 Abdominal sternites light brown ( Figs 1a, b )................................................................ 2

- Abdominal sternites entirely metallic ( Fig. 1c ).............................................................. 4

2 Meso-, metasternum, metepisternum and abdomen light brown ( Fig 1a ).......................................... 3

- Mesosternum, all episterna and sides of metasternum metallic, central part of metasternum and abdomen light brown ( Fig. 1b )................................................................................ P. (S.) kapuka sp. nov.

3 Striae on vertex strong; pronotum gently transversally rugate; elytra coarsely punctate; yellow apical band of elytra reaching suture along margins ( Fig. 11 ). Head as wide as elytra (mean EW/HW = 1.0)................... P. (S.) inopinata sp. nov.

- Striae on vertex shallow, pronotum smooth; elytra shiny, sparsely and weakly punctate; yellow apical band of elytra rounded ( Fig. 10 ). Head wider than elytra (EW/HW = 0.89).................................... P. (S.) incerata Rivalier, 1964

4 Punctures on elytral surface rounded at base and behind humeri; transverse, narrow and often confluent on the disc; becoming squamiform in posterior part............................................................................ 5

- Elytra uniformly sculptured laterally and on the disc. Punctures deeply impressed and rounded, slightly transverse in apical part; never squamiform. Body slender, upper side mate, blue or coppery red ( Figs 4–5 )........ P. (S.) coerulea Horn, 1920

5 Aedeagus elongate, narrow in basal third and gradually widened on dorsal side to the middle ( Figs 7 , 9 )................ 6

- Aedeagus short, abruptly widened immediately from the base ( Figs 3 , 13 )........................................ 7

6 Dorsal side of body shiny, with bright copper and green reflections. Anterior half of pronotum smooth and shiny ( Fig. 6 ). Body relatively robust. Apex of aedeagus narrow, with rounded hook spiral sclerite of endophallus with two full coils at base ( Fig. 7 )................................................................................. P. (S.) eureka sp. nov.

- Dorsal side of body dull, dark copper. Pronotum evenly, transversely rugate throughout ( Fig. 8 ). Apex of aedeagus slightly wider, with down turned tip; spiral sclerite of endophallus wide, with two and half full coils at base ( Fig. 9 )...................................................................................... P. (S.) heteromallicollis Horn, 1909

7 Pronotum dull, densely transversely rugate. Disc of elytra with coarse transverse punctures, becoming confluent in apical half. Aedeagus short and robust; apex short and wide, with thick hook; spiral sclerite of endophallus wide, with one and half full coils at base ( Fig. 13 ).................................................................. P. (S.) sciaky sp. nov.

- Pronotum shiny, moderately transversally rugate. Apex of aedeagus not thickened, with more or less pointed hook-shaped denticle; spiral sclerite of endophallus long and narrow, with three full coils at base ( Figs 2–3 ).................................................................................................... P. (S.) aeneoparva Horn, 1934

Figures

FIGURE 1. Ventral habitus of Prothyma (Symplecthyma). a) P. inopinata sp. nov.; b) P. kapuka sp. nov.; c) P. aeneoparva Horn, 1934. Scale = 5 mm.

FIGURE 2. Prothyma (Symplecthyma) aeneoparva Horn, 1934. a–f: Dorsal habitus. Scale = 5 mm. a) Female, Leyte. b) P. nitida Rivalier 1964 syn. nov., Type, male. c) P. boholensis Dheurle 2012 syn. nov., Holotype, female. d) Male from Cawa- Cawa, Mindanao. e) Male, Park Tayabas, Luzon. f) Lectotype, male, Samar. g) Labrum, Male. h) Labrum, Female. Scale = 1 mm.

FIGURE 3. Aedeagus of Prothyma (Symplecthyma) aeneoparva Horn, 1934, left lateral view. a) Syntype, Biliran; b) Lope DeVega, Samar, lateral and ventral view; c) Syntype, Samar; d) Hinabangan, Samar; e) P. nitida Rivalier 1964 syn. nov., Type, drawing from (Rivalier, 1964); f) Segod, Leyte; g) Park Tayabas, Luzon; h) Surigao, Mindanao; i) Wao, Mindanao; j) Sibagat; k) Cawa-Cawa, Mindanao; l) Gutalac, Mindanao; m) Cabanglasan, Mindanao; n) Esperanza, Mindanao. Scale = 1 mm.

FIGURE 4. Prothyma (Symplecthyma) coerulea Horn, 1920. a) Dorsal habitus Lectotype, male. b) Dorsal habitus Holotype P. bella Dheurle & Colas, 2018, syn. nov. = P. coerulea opacosericea Horn, 1929 comb. nov.; c) Dorsal habitus Lectotype, male P. coerulea opacosericea Horn, 1929 comb. nov. Scale = 5 mm. d–e Labrum P. coerulea s.str. ST. Scale = 1 mm.: d) Male, e) Female. f–h. Aedeagus. Scale = 1 mm. f) P. coerulea Lectotype. g) Lectotype, male P. coerulea opacosericea Horn, 1929 comb. nov.. h) Holotype P. bella Dheurle & Colas, 2018, syn. nov. = P. coerulea opacosericea Horn, 1929 comb. nov.

FIGURE 5. Head of Prothyma (Symplecthyma) coerulea Horn, 1920. a) P. coerulea s.str. b) P. bella Dheurle & Colas, 2018, syn. nov., c) P. coerulea opacosericea Horn, 1929. Scale = 1 mm.

FIGURE 6. Prothyma (Symplecthyma) eureka sp. nov. a) Dorsal habitus. Holotype male; b) Elytra, paratype, female. Scale = 5 mm. c) Labrum, female; d) Labrum, male. Scale = 1 mm.

FIGURE 7. Aedeagus of Prothyma (Symplecthyma) eureka sp. nov. a–b) Mt. Apo; c–d) Mt. Kitanglad; e–f) San Miguel; g–i) Agusan del Sur, Esperanza. Scale = 1 mm.

FIGURE 8. Prothyma (Symplecthyma) heteromallicollis Horn, 1909. a) Dorsal habitus, Holotype, male. b) Elytra, female. Scale = 5 mm. c) Head and labrum, male; d) Head and labrum, female. Scale = 1 mm.

FIGURE 9. Aedeagus of Prothyma (Symplecthyma) heteromallicollis Horn, 1909. a–b) Holotype; c–e) Luzon, Cagayan, St. Ana; f) Mindanao, Luagon; g–h) Mindanao, Nabunturan; i–j) Mindanao, Davao de Oro. Scale = 1 mm.

FIGURE 10. Prothyma (Symplecthyma) incerata Rivalier, 1964. a) Dorsal habitus holotype, male. Scale = 5 mm. b) Aedeagus. c) Endophallus. Drawings from (Rivalier, 1964).

FIGURE 11. Prothyma (Symplecthyma) inopinata sp. nov. a) Dorsal habitus holotype female. b) Dorsal habitus paratype female. Scale = 5 mm. c) Labrum. Scale = 1 mm.

FIGURE 13. Prothyma (Symplecthyma) sciakyi sp. nov. a) Dorsal habitus, Holotype, male. Scale = 5 mm. b) Left elytron, paratype, female. Scale = 5 mm. c) Labrum, male; d) Head, female; e) Aedeagus, left lateral view, Governor Generoso; f) Aedeagus, left lateral view, San Francisco Scale = 1 mm.