Ypopteron nicola, Archibald & Gu & Mathewes, 2022

Archibald, S. Bruce, Gu, Jun-Jie & Mathewes, Rolf W., 2022, The Palaeorehniidae (Orthoptera, Ensifera, “ Zeuneropterinae ”), and new taxa from the early Eocene Okanagan Highlands, western North America, Zootaxa 5100 (4), pp. 559-572 : 566-567

Ypopteron nicola new species

( Fig. 4 )

Haglidae , Prophalangopsinae sp.: Archibald and Mathewes 2000: 1443, Fig. 3 .

Diagnosis. As for genus.

Description. Holotype tegmen. As in diagnosis and: about 6.5 cm long as preserved (small portion of basalmost region missing); membrane apparently infuscate throughout; RA, RP branch distal branching of MA, MP; RP branching eight times, one branch branched again near margin; MA, MP branch at level of CuA joining CuPaα, distally parallel with branches of RP; CuA+CuPaα with five branches preserved, probably total; no stridulatory file detected on CuPb; simple crossveins throughout proximal posterior portion; zigzag intercalated crossveins forming pentagons in apical portion of overlapping wing between branches of RP.

Type specimen. Holotype. Q-1010 , part (no counterpart), two or more wings, two almost perfectly overlapping. The anterior regions (RA, stem of RP, ScA, SCP and precostal region) are missing or covered by portions of one or more wing(s); labelled Ypopteron nicola Archibald, Gu, and Mathewes , holotype; collected by Jurgen Mathewes, 1989; deposited in the collections of Simon Fraser University.

Locality and age. Quilchena, British Columbia; Coldwater Beds; mid Ypresian, 51.5 ± 0.4 Ma. old.

Etymology. The specific epithet refers to the Nicola Valley, where the Quilchena locality is located.

Figures

FIGURE 3. Diagnostic character states of the wing of Palaeorehniidae (see text); comparative Prophalangopsidae (A) and Palaeorehniidae (B, C): A, Aboilus tigris Gorochov, female, redrawn from Gorochov (1996, Fig. 2c); B, Republicopteron douseae specimen SR 21-005-001; C, tegmen of Albertoilus cervirufi, redrawn from Kevan and Wighton, with distal fragment positioned as in their drawing (1983, Fig. 1b). Scale bar for B, see Fig. 7; for C, see original; original of A has no scale bar.

FIGURE 4. Ypopteron nicola, new genus and species, holotype Q-1010, Quilchena: A, photograph; B, drawing. Scale bar = 1 cm.