Condyloderes storchi Higgins, 2004 in Martorelli & Higgins, 2004

Neuhaus, Birger, Zotto, Matteo Dal, Yamasaki, Hiroshi & Higgins, Robert P., 2019, Revision of Condyloderes (Kinorhyncha, Cyclorhagida) including description of Condyloderes shirleyi sp. nov., Zootaxa 4561 (1), pp. 1-91 : 56

Re-description of Condyloderes storchi Higgins, 2004 in Martorelli & Higgins, 2004

( Figs 29–31 ; Tables 1, 2, 7)

Condyloderes storchi Higgins, 2004 in Martorelli & Higgins, 2004 : pp. 86–88, figs 2–5, tables 1, 2.

Condyloderes storchi — Sørensen & Pardos 2008: pp. 53, 54 [species erroneously cited as C. storchi Martorelli & Higgins, 2004 ].— Shirley 2009: p. 1129 [species erroneously cited as C. storchi Martorelli & Higgins, 2004 ].— Neuhaus et al. 2010: p. 468.— Sørensen et al. 2010b: pp. 235, 241 [species erroneously cited as C. storchi Martorelli & Higgins, 2004 ].— Neuhaus 2013, pp. 297, 332, tables 6, 12.— Adrianov & Maiorova 2016: pp. 11, 18 [species erroneously cited as C. storchi Martorelli & Higgins, 2004 ].

Diagnosis. Neck placids with condyles in one basal row in narrower placids and three rows in broader placids; narrower placids with one basal condyle; broader placids with two condyles each in apical, intermediate, and basal row; midventral placid with three apical and basal condyles and two in intermediate row; acicular spine middorsally on segments 1–10, midterminally on segment 11, lateroventrally on segments 1–9, and laterodorsally on segment 10; cuspidate spine ventrolaterally on segments 5 and 9, lateral accessorily on segments 1, 4 and 8, and middorsally on segments 5 and 7; type-5 (?) sensory spot ventromedially on segment 11; type-6 sensory spot ventromedially on segments 1 (slightly more paraventrally) and 2–9, lateroventrally on segment 10, sublaterally on segments 1–3, 5–7 and 9, midlaterally on segments 1 and 2, laterodorsally on segments 1–9, subdorsally on segment 10, and paradorsally on segments 1 (almost subdorsally), 2, 4–6, 8 and 9.

Material examined. Holotypic male (USNM 1025879) mounted laterally ( Fig. 1 ; Table 1).

Figures

FIGURE 1. Map of locations of all known species of Condyloderes identified to species level; see Table 1 for location data. Filled symbols indicate specimens studied in this article, open symbols refer to specimens not investigated here.

FIGURE 29. Condyloderes storchi. A, B. Habitus of holotypic male (USNM 1025879) from right (A) and ventrolateral-left side (B). Cuspidate spines highlighted by grey colour. Scale bar 100 µm, valid for A and B. Abbreviations: ac, acicular spine; cu, cuspidate spine; lf10, laterodorsal spine of segment 10; ltas, lateral terminal accessory spine; md7, middorsal spines of segment 7; mts, midterminal spine; ne, protonephridial openings; ssp3, type-3 sensory spot.

FIGURE 30. Condyloderes storchi, holotypic male (USNM 1025879) from right side, DIC. A. Head and neck with placids; note condyles of placids. Inset with inner oral styles. B. Neck, placids and basal plate of trichoscalids. C. Segments 1–3; extended focus image of four images. D. Dorsal part of segments 5–7. E. Segments 4–6; extended focus image of two images. F. Segments 7–9; extended focus image of five images. Notice middorsal cuspidate spine on segments 5 and/or 7 in D–F. Arrowheads in C–F mark type-6 sensory spots. Scale bar in C, E, and F 50 µm. Scale bar in D 50 µm, valid for A, B, and D. Abbreviations: ac6, acicular spine of segment 6; bpl, broader placid; co, condyle of placid; cu, cuspidate spine; dv, attachment area of dorsoventral muscle; ios, inner oral style; mc, mouth cone; md3, middorsal spine of segment 3; ne, protonephridial openings; npl, narrower placid; s1, segment 1; sc01, spinoscalid of ring 01; sf, secondary fringe; tr, trichoscalid or its basal plate.

FIGURE 31. Condyloderes storchi, holotypic male (USNM 1025879) from ventrolateral-left (A–E) and right (F) side, DIC. A. Placids; note appendages of trichoscalids. B. Segments 1–4; arrows mark lack of free flap ventromedially on segment 1. C. Segments 5–7; extended focus image of two images. D. Segments 8 and 9; extended focus image of two images. E, F. Segments 10 and 11; arrow in E points to type-3 sensory spot. Arrowheads in B–F mark type-6 sensory spots. Scale bar in B 50 µm, valid for A, B, and E. Scale bar in C, D, and F 50 µm. Abbreviations: co, condyle of placid; cu, cuspidate spine; dv, attachment area of dorsoventral muscle; fa, appendage at trichoscalid; ld10, laterodorsal spine of segment 10; ltas, lateral terminal accessory spine; lv2, lateroventral spine of segment 2; md10, middorsal spine of segment 10; ne, protonephridial openings; sf, secondary fringe; st, sternal plate; tp, tergal plate; tr, trichoscalid.