Raveniola adjarica, Zonstein & Kunt & Yağmur, 2018

Zonstein, Sergei, Kunt, Kadir B. & Yağmur, Ersen A., 2018, A revision of the spider genus Raveniola (Araneae, Nemesiidae). I. Species from Western Asia, European Journal of Taxonomy 399, pp. 1-93 : 8-10

publication ID	https://doi.org/ 10.5852/ejt.2018.399
publication LSID	lsid:zoobank.org:pub:E836E138-D6E2-4F62-B4B3-CE2E073F2B24
DOI	https://doi.org/10.5281/zenodo.5980277
persistent identifier	https://treatment.plazi.org/id/BCF871CB-E0C5-4A75-B376-EA83C72B04A5
taxon LSID	lsid:zoobank.org:act:BCF871CB-E0C5-4A75-B376-EA83C72B04A5
treatment provided by	Plazi
scientific name	Raveniola adjarica
status	sp. nov.

Treatment

Raveniola adjarica View in CoL sp. nov.

urn:lsid:zoobank.org:act:BCF871CB-E0C5-4A75-B376-EA83C72B04A5

Figs 43–44 View Figs43–51 , 84 View Figs 84–89 , 102 View Figs 102–116 , 117 View Figs 117–128 , 129 View Figs129–132 , 149 View Figs 149–152 , 171–173 View Figs 171–179 , 198 View Figs 198–203 , 219 View Figure 219

Brachythele pontica Spassky, 1937: 368 (♀, part; misidentified).

Diagnosis

Different from other species in the same group by having a relatively short embolus with a rounded submedian keel ( Figs 171–173 View Figs 171–179 ; cf. Figs 174–179 View Figs 171–179 ) and long, asymmetrical spermathecae with median receptacles being considerably longer than laterals ( Fig. 198 View Figs 198–203 , cf. Figs 199–203 View Figs 198–203 ).

Etymology

The specific epithet, a Latinized adjective, is derived from Adjaria (also Adjara or Adzharia), the historical region and administrative province of Georgia where the species was found.

Material examined

Holotype GEORGIA: ♂, South slope of Meskheti Mts, Kintrishi Valley, 3 km NE of Khino , 900–950 m, 41°43.8′ N, 42°03.5′ E, 7 Sep. 1989, S.L. Zonstein leg. ( TAU) GoogleMaps .

Paratypes (1 ♂, 3 ♀♀) GEORGIA: 2 ♀♀, same collection data as for holotype, but 7–8 Sep. 1989 ( TAU) GoogleMaps ; 1 ♂, 1 ♀, Kintrishi Valley, 0.5 km W of Zerabozeli , 450 m, 41°44.0′ N, 41°59.2′ E, 15 Sep. 1989, S.L. Zonstein leg. ( TAU). GoogleMaps

Additional material (2 ♀♀, 1 juv.)

GEORGIA: 1 ♀, Batumi, 1909, S.A. Spassky leg. ( ZISP, paralectotype of Brachythele pontica ); 1 ♀, same locality, no other collecting data ( MNHN 10338 ); 1 juv., surroundings of Batumi, Mt Feria , 250 m, 9 Feb. 1982, D.V. Logunov leg. ( TAU).

Description

Male (holotype)

HABITUS. See Fig. 43. View Figs43–51

MEASUREMENTS. TBL 13.10, CL 5.43, CW 4.88, LL 0.47, LW 1.03, SL 2.52, SW 2.38.

COLOUR. Carapace brownish rufous with slightly lighter yellowish rufous pattern covering foveal area and protruding into few weak radial rays on darker background; eye tubercle with central and two symmetrical lateral brownish black spots surrounding AMEs and lateral eyes respectively; chelicerae and first pair of legs reddish brown; palps and legs II–IV brownish rufous; sternum, labium and maxillae light yellowish rufous; abdomen dorsally light greyish brown, with weak and darker brownish pattern represented by median lanceolate patch fused and crossed with few transverse fasciae; spinnerets and ventral surface of abdomen light yellowish grey.

PROSOMA. Carapace and chelicerae as shown in Fig. 66 View Figs 59–67 . Clypeus and eye group as in Fig. 102 View Figs 102–116 . Eye diameters and interdistances: AME 0.15(0.20), ALE 0.23, PLE 0.15, PME 0.13, AME–AME 0.11(0.06), ALE–AME 0.10(0.08), ALE–PLE 0.08, PLE–PME 0.04, PME–PME 0.40. Each cheliceral furrow with 10 promarginal teeth and 5 mesobasal denticles. Maxillae with 7–8 cuspules each.

LEGS. Tibia and metatarsus I as in Fig. 129 View Figs129–132 . Scopula: in distal 1/3 of metatarsi I–II, entire on tarsi I– II, vestigial and widely divided by setae on tarsus III, absent on tarsus IV. Trichobothria: 2 rows of 9–11 per row on tibiae, 10–15 on metatarsi, 10–12 on tarsi, 9 on cymbium. Paired claws of tarsi I–II: outer and inner margins with 7–8 teeth each; claws of tarsi III–IV: outer and inner margins with 8–10 teeth each.

SPINATION. Palp: femur d1–1–1–1(2), pd1, rd1; patella p1; tibia d1, p1–1–1, r1–1, v2–2–2; tarsus d4. Leg I: femur d1–1–1–1, pd1–1–1, rd1; tibia p1–1(0), pv1–1, rv1–1–m–m; metatarsus v0–0–3. Leg II: femur d1–1–1–1, pd1–1–1; patella p1; tibia p1–1–1, v2–2–3; metatarsus p1, v1–2–3. Leg III: femur d1–1–1–1, pd1–1(0)–1, rd1–1(0) –1; patella p1, r1; tibia d1–1, p1–1, r1–1–1, v2–2–3; metatarsus d1–1, p1–1–1, r1–1–1, v2–3–3. Leg IV: femur d1–1–1–1–1(0), pd1, rd1(0)–1; patella r1; tibia d1–1, p1–1, r1–1, v2– 2–3; metatarsus d1–1, p1–1–1–1, r1–1(0)–2–1–2, v1–2–2–1–3–3. Patella I and tarsi I–IV (except right tarsus III carrying 1 retrolateral spine) aspinose.

PALP. Tibia, cymbium and palpal organ as shown in Fig. 149 View Figs 149–152 . Wide and flattened basal part of embolus provided with subapical keel extends into twisted and narrow apical part of embolus ( Figs 171–173 View Figs 171–179 ).

SPINNERETS. PMS: length 0.75; diameter 0.28. PLS: maximal diameter 0.65; length of basal, medial and apical segments 1.22, 0.85, 1.23; total length 3.30; apical segment digitiform.

LEG MEASUREMENTS. ♂(♀).

	Femur	Patella	Tibia	Metatarsus	Tarsus	Total
Palp	2.95 (3.07)	1.67 (1.95)	2.35 (2.20)	–	1.02 (2.12)	7.99 (9.34)
Leg I	4.55 (4.07)	2.60 (2.57)	3.57 (2.95)	3.28 (2.43)	2.01 (2.12)	16.01 (13.77)
Leg II	4.02 (3.55)	2.25 (2.23)	3.03 (2.72)	2.85 (2.40)	1.78 (1.73)	14.23 (12.63)
Leg III	3.17 (3.03)	1.72 (1.90)	2.46 (2.17)	3.22 (2.75)	1.79 (1.73)	12.36 (11.58)
Leg IV	4.47 (4.10)	2.20 (2.25)	3.43 (3.23)	4.15 (3.88)	2.02 (2.07)	16.27 (15.53)

Female (paratype, from Khino)

HABITUS. See Fig. 44. View Figs43–51

MEASUREMENTS. TBL 15.90, CL 6.17, CW 4.48, LL 0.62, LW 1.33, SL 2.85, SW 2.60.

COLOUR. As in male, but with lighter, quite indistinct radial pattern on carapace and with lighter (yellowish rufous) clypeus.

PROSOMA. Carapace and chelicerae as shown in Fig. 84 View Figs 84–89 . Clypeus and eye group as in Fig. 117 View Figs 117–128 . Eye diameters and interdistances: AME 0.14(0.19), ALE 0.22, PLE 0.13, PME 0.13, AME–AME 0.14(0.08), ALE–AME 0.15(0.12), ALE–PLE 0.07, PLE–PME 0.07, PME–PME 0.51. Each cheliceral furrow with 10 promarginal teeth and 4–5 mesobasal denticles. Maxillae with 5–7 cuspules each.

LEGS. Scopula: distal on metatarsi I–II, entire on tarsus I and on palpal tarsus, narrowly divided by setae on tarsus II, elsewhere absent. Trichobothria: 2 rows of 11–14 each on tibiae, 11–14 on metatarsi, 10–12 on tarsi, 8 on palpal tarsus. Paired claws: outer and inner margins with 6–8 teeth each, palpal claw with 4 teeth on promargin.

SPINATION. Palp: femur d1+2b, pd1; tibia v2–2–3; tarsus v2–2–3. Leg I: femur d1+3b, pd1; tibia v2– 2–2; metatarsus v2–2–3. Leg II: femur d1+3b, pd1; tibia p1–1, v2–2–3; metatarsus p1, v2–2–3. Leg III: femur d1+3(4)b, pd2b, rd1–1; patella r1; tibia p1–1, r1–1, v2–2–3; metatarsus p1–1, r1–1, v2–2–3. Leg IV: femur d1+3b, pd2b, rd1; patella r1; tibia p1–1, r1–1, v2–2–3; metatarsus p1(0)–1–1, r1–1–1, v2–1(0)–2–3. Patellae I–II, and palpal patella aspinose.

SPERMATHECAE. Broad and asymmetrical, with widely spaced receptacles ( Fig. 198 View Figs 198–203 ).

SPINNERETS. PMS: length 0.36; diameter 0.26. PLS: maximal diameter 0.55; length of basal, medial and apical segments 0.70, 0.37, 0.65; total length 1.72; apical segment shortly digitiform.

Variation

Carapace length in males varies from 5.15 to 5.38, in females from 4.55 to 6.20. No variation in the structure of the male palpal organ has been observed. Female vulvas look surprisingly uniform and similar to the described type, even that in the conspecific female MNHN 10338 from Batumi, the known locality that is farthest from the type locality.

Ecology

The spiders collected from the Kintrishi area were found hiding in soil cavities under stones in chestnut and spruce mountain forests dominated by Castanea sativa Mill. and Picea orientalis L. (Link.), respectively.

Distribution

Known from the western part of the Caucasus Minor (Georgia: Adzharia). See Fig. 219 View Figure 219 .