Raveniola nana, Zonstein & Kunt & Yağmur, 2018

Zonstein, Sergei, Kunt, Kadir B. & Yağmur, Ersen A., 2018, A revision of the spider genus Raveniola (Araneae, Nemesiidae). I. Species from Western Asia, European Journal of Taxonomy 399, pp. 1-93 : 23-24

Raveniola nana sp. nov.

urn:lsid:zoobank.org:act:C3C6CE26-9C99-4A76-9A63-46779FEFF1CA

Figs 89 , 122 , 204 , 221

Diagnosis

The structure of the spermathecae of Raveniola nana sp. nov. resembles that of R. turcica sp. nov. (the female of R. birecikensis sp. nov. is unknown), but differs in having the median and lateral receptacles situated considerably closer to each other ( Fig. 204 ; cf. Fig. 205 ). Additionally, R. nana sp. nov. is considerably smaller and has a paler coloration than both the latter species.

Etymology

The specific epithet (a variable adjective from Ancient Greek νᾶνος, meaning ‘dwarf’) refers to the very small size of this species.

Material examined

Holotype TURKEY: ♀, Hatay Province, Hassa District, Küreci Village , 36°42′36.89″ N, 36°27′14.44″ E, 499 m, 15 Nov. 2010, Anlaş and Yağmur leg. ( AUZM).

Description

Female (holotype)

MEASUREMENTS. TBL 10.97, CL 3.69, CW 2.82, LL 0.29, LW 0.65, SL 1.84, SW 1.62.

COLOUR. Most parts of body pale brownish; eye tubercle with partially fused dark brown rings surrounding AMEs and lateral eyes; chelicerae light red; abdomen uniformly light yellowish grey without dorsal pattern, spinnerets pale yellowish grey.

PROSOMA. Carapace and chelicerae as shown in Fig. 89 . Clypeus and eye group as in Fig. 122 . Eye diameters and interdistances: AME 0.09(0.12), ALE 0.13, PLE 0.10, PME 0.08, AME–AME 0.08(0.06), ALE–AME 0.05(0.04), ALE–PLE 0.05, PLE–PME 0.02, PME–PME 0.23. Each cheliceral furrow with 8 promarginal teeth and 7 mesobasal denticles. Maxillae each with 5–7 cuspules along heel.

LEGS. Scopula: distal on metatarsi I–II, entire on tarsus I and palpal tarsus, widely divided on tarsus II, elsewhere absent. Trichobothria: 2 rows of 8–9 each on tibiae, 9–10 on metatarsi, 10–11 on tarsi, 8 on palpal tarsus. Paired tarsal claws with 7–10 teeth on each margin. Palpal claw with 6 teeth on inner margin.

LEG MEASUREMENTS.

	Femur	Patella	Tibia	Metatarsus	Tarsus	Total
Palp	1.82	1.24	1.26	–	1.15	5.47
Leg I	2.24	1.53	1.85	1.50	1.12	8.24
Leg II	2.17	1.35	1.65	1.45	1.16	7.78
Leg III	2.07	1.15	1.45	1.95	1.19	7.81
Leg IV	2.58	1.37	2.35	2.83	1.36	10.49

SPINATION. Femora I–IV with one basodorsal spine and 3 dorsal bristles alongside midline; palpal femur dorsally with 3 bristles; palpal patella and patellae I–II aspinose. Palp: femur pd0–0–1; tibia v1–1–2; tarsus v2–0–0. Leg I: femur pd0–0–1; tibia v1–1–2; metatarsus v1–2–2. Leg II: femur pd0–0–1; tibia p0–0–1, v1–2–2; metatarsus v1(0)–1–2. Leg III: femur pd1–1–1, rd1–1–1; patella p1, r1; tibia d1–1(0)– 0, p1–1–1, r1–1–1, v2–2–3; metatarsus pd0–1–0, p1–1–1, r1–1–1, v2–2–3. Leg IV: femur pd0–0–1, rd0–0–1; patella r1; tibia d1–0–0, p1–1–1, r1(0)–1–1, v2–2–3; metatarsus d0–1–1–0, p1–1–1–1, r1–1– 1–1, v2–2–3.

SPERMATHECAE. Moderately broad with receptacles close to each other ( Fig. 204 ).

SPINNERETS. PMS: length 0.36; diameter 0.16. PLS: maximal diameter 0.31; length of basal, medial and apical segments 0.55, 0.35, 0.34; total length 1.24; apical segment shortly digitiform.

Ecology

The holotypic female was found in open forest under a stone.

Distribution

South-Eastern Turkey ( Fig. 221 ).

Note

There is a possibility that the holotype of Raveniola nana sp. nov. is the unknown female of the allopatric R. biresekensis sp. nov. However, both these species differ in size and colouration ( R. nana sp. nov. is noticeably smaller and considerably paler than the latter species), as well as in the relative size of AMEs and PMEs (larger vs smaller, respectively). We thus prefer to consider these names as belonging to different species, until the currently missing males of R. nana sp. nov. and the unknown females of R. biresekensis sp. nov. are collected and identified.

Figures

Figs 84–89. Raveniola spp., ♀♀. Carapace and chelicerae, dorsal views. — 84. R. adjarica sp. nov. (paratype). 85. R. dunini sp. nov. (paratype). 86. R. hyrcanica Dunin, 1988 (paratype). 87. R. pontica (Spassky, 1937) (Lazarevskoe). 88. R. sinani sp. nov. (holotype). 89. R. nana sp. nov. (holotype). Scale bars: 1.0 mm.

Figs 117–128. Raveniola spp., ♀♀. Eyes and clypeus, dorsal views. — 117. R. adjarica sp. nov. (paratype).118. R.dunini sp. nov.(paratype).119.R.hyrcanica Dunin, 1988(paratype).120. R.pontica (Spassky, 1937) (Lazarevskoe). 121. R. sinani sp. nov. (holotype). 122. R. nana sp. nov. (holotype). 123. R. turcica sp. nov. (paratype). 124. R. micropa (Ausserer, 1871) (holotype). 125. Same species as preceding (Uludağ). 126. R. zaitzevi (Charitonov, 1948) (Lagodekhi). 127. R. vonwicki Zonstein, 2000 (paratype). 128. R. niedermeyeri (Brignoli, 1972) (paratype). Scale bar: 0.5 mm.

Figs 204–212. Raveniola spp., ♀♀. Vulva, dorsal views. — 204. R. nana sp. nov. (holotype). 205. R. turcica sp. nov. (paratype). 206. R. micropa (Ausserer, 1871) (Uludağ). 207. Same aspecies as preceding (holotype). 208. R. zaitzevi (Charitonov, 1948) (Lagodekhi). 209. Same species as preceding (Pirkuli). 210. R. vonwicki Zonstein, 2000 (paratype). 211. R. niedermeyeri (Brignoli, 1972) (paratype, Gorgan). 212. Same species as preceding (Mazandaran). Scale bars: 0.25 mm.

Figure221. Localitiesof Raveniolaspp.inTurkey:R.anadolu sp.nov.(),R.arthuriKunt&Yağmur, 2010(),R.birecikensissp.nov.(),R.micropa(Ausserer,1871)(),R.nanasp.nov.(),R.sinani sp. nov.(), and R. turcica sp. nov. ().Abbreviation:AM =Armenia.