Raveniola marusiki, Zonstein & Kunt & Yağmur, 2018

Zonstein, Sergei, Kunt, Kadir B. & Yağmur, Ersen A., 2018, A revision of the spider genus Raveniola (Araneae, Nemesiidae). I. Species from Western Asia, European Journal of Taxonomy 399, pp. 1-93 : 37-38

Raveniola marusiki sp. nov.

urn:lsid:zoobank.org:act:C48F164E-60BD-4454-A24D-FC7459D30BDF

Figs 81 , 114 , 142 , 162 , 191–192

Diagnosis

In the shape of the distally spiraled embolus, the holotype of Raveniola marusiki sp. nov. resembles males of R. vonwicki and R. zaitzevi , but differ from them, as well as from other species of the same group, by the unique shape of its pectinate embolic keel ( Fig. 191 ).

Etymology

The specific epithet is given in honour of our friend and colleague Yuri Marusik for his immense contribution to the study of the spider fauna of Turkey and Iran.

Material examined

Holotype IRAN: ♂, Golestan Province, Elburs Mts, Germabdasht Valley 15–20 km SE of Gorgan , 36°48′ N, 54°34′ E, 500–1000 m, 7–9 Oct. 1903, N.A. Zarudny leg. ( ZMMU).

Description

Male (holotype)

MEASUREMENTS. TL 14.50, CL 5.64, CW 4.96, LL 0.49, LW 0.97, SL 2.58, SW 2.53.

COLOUR. Carapace, palps and legs dorsally brownish rufous; eye tubercle anteriorly brownish black, chelicerae reddish brown; sternum, labium, maxillae, palps and legs ventrally yellowish brown; abdomen dorsally brown, with darker pattern consisting of partially fused dark brown irregular spots; ventral surface of abdomen light greyish brown.

PROSOMA. Carapace and chelicerae as shown in Fig. 81 . Clypeus and eye group as in Fig. 114 . Eye diameters and interdistances: AME 0.15(0.19), ALE 0.25, PLE 0.18, PME 0.13, AME–AME 0.10(0.05), ALE–AME 0.08(0.06), ALE–PLE 0.11, PLE–PME 0.04, PME–PME 0.37. Each cheliceral furrow with 9 promarginal teeth and 6 mesobasal denticles. Maxillae: each with 9 relatively small cuspules.

LEGS. Tibia and metatarsus I as shown in Fig. 142 . Scopula: distal on metatarsi I–II, entire on tarsi I– II; absent on tarsi III–IV. Trichobothria: 2 rows of 9–10 each on tibiae, 10–14 on metatarsi, 10–12 on tarsi, 7–8 on cymbium. Paired claws with 8–11 teeth on each margin.

LEG MEASUREMENTS.

	Femur	Patella	Tibia	Metatarsus	Tarsus	Total
Palp	3.01	1.72	2.37	–	0.95	8.05
I	4.59	2.76	3.67	3.36	2.18	16.56
II	3.90	2.14	3.03	3.24	2.00	14.31
III	3.76	1.88	2.51	3.57	2.01	13.73
IV	4.75	2.25	3.70	5.12	2.28	18.10

SPINATION. Patella I aspinose. Palp: femur d0–1–1–1, pd1; patella p1; tibia d1–1–0, p2–2–2, r1–1, pv0–0–1, rv1–1–0; cymbium d6. Leg I: femur d1–1–1–1, pd1–1–1, rd0–0–1; tibia p1–1–0, pv v1–0–1–0; rv0–1–0– m–m; metatarsus v0–0–2. Leg II: femur d1–1–1–1, pd0–1–1; tibia p1–1–1, v2–2–2; metatarsus p0–1–0; v2–2–2. Leg III: femur d1–1–1–1, pd1–1–1, rd1–1–1; patella p1–1(0), r1; tibia d1–1–0, p1–1–1, r1–1– 1, v2–2–3; metatarsus d1–0–1, p1–1–1, r1–1–1, v2–2–3. Leg IV: femur d1–1–1–1, pd1–1–1, rd0–0–1; patella r1; tibia d1–1–0, p1–1–1, r1–1–1, v2–2–3; metatarsus d1–0–1–0, p2–2–1–1, r1–1–1–1, v2–2–2–3.

PALP. Tibia, cymbium and palpal organ as shown in Fig. 162 . Short pectinate embolic keel consists of two large and a few smaller teeth; embolus gently corkscrew-shaped, with proximal bend located submedially ( Figs 191–192 ).

SPINNERETS. PMS: length 0.48; diameter 0.21. PLS: maximal diameter 0.45; length of basal, medial and apical segments 0.71, 0.55, 0.58; total length 1.84; apical segment shortly digitiform.

Female

Unknown.

Ecology

Judging from the label data, the holotypic male was collected in the broad-leaved forest zone of Alborz Mts. Other details are unknown.

Distribution

Known only from the type locality.

Figures

Figs 79–83. Raveniola spp., ♂♂. Carapace and chelicerae, dorsal views. — 79. R. mazandaranica Marusik,Zamani&Mirshamsi,2014(holotype).80.R.zaitzevi(Charitonov,1948)(Car).81.R.marusiki sp. nov. (holotype). 82. R. vonwicki (Zonstein, 2000) (holotype). 83. R. niedermeyeri (Brignoli, 1972) (holotype). Scale bars: 1.0 mm.

Figs 102–116. Raveniola spp., ♂♂. Eyes and clypeus, dorsal views. — 102. R. adjarica sp. nov. (holotype).103. R.duninisp.nov.(holotype).104. R.hyrcanicaDunin, 1988(holotype).105.R.pontica (Spassky, 1937) (Lazarevskoe). 106.R.birecikensis sp. nov.(holotype).107. Samespecies as preceding (dwarf conspecific male?). 108. R. anadolu sp. nov. (holotype). 109. R. arthuri Kunt & Yağmur, 2010 (holotype). 110. R. turcica sp. nov. (holotype). 111. R. micropa (Ausserer, 1871) (Uludağ). 112. R. mazandaranica Marusik, Zamani & Mirshamsi, 2014 (holotype). 113. R. zaitzevi (Charitonov, 1948) (Car). 114. R. marusiki sp. nov. (holotype). 115. R. vonwicki (Zonstein, 2000) (holotype). 116. R. niedermeyeri (Brignoli, 1972) (holotype). Scalebar: 0.5 mm.

Figs 141–144. Raveniola spp., ♂♂. Tibia and metatarsus I, retrolateral views. — 141. R. zaitzevi (Charitonov, 1948) (Car). 142. R. marusiki sp. nov. (holotype). 143. R. vonwicki Zonstein, 2000 (holotype). 144. R. niedermeyeri (Brignoli, 1972) (holotype). Scale bars: 1.0 mm.

Figs 161–164. Raveniola spp., ♂♂. Palpal tibia, cymbium and palpal organ, retrolateral views. — 161.R.zaitzevi(Charitonov,1948)(Car).162.R.marusikisp.nov.(holotype).163.R.vonwickiZonstein, 2000 (holotype). 164. R. niedermeyeri (Brignoli, 1972) (holotype). Scale bars: 1.0 mm.

Figs 189–197. Raveniola niedermeyeri species group, holotypic (189–192, 195–197) and conspecific (193–194) ♂♂. Details of palpal organ; retrolateral (189, 191, 194–195, 197) and ventral (190, 192–193, 196) views. — 189–190. R. mazandaranica Marusik, Zamani & Mirshamsi, 2014. 191, 192.R.marusikisp.nov.193–194. R.zaitzevi(Charitonov, 1948).195–196. R.vonwickiZonstein,2000. 197. R. niedermeyeri (Brignoli, 1972). White arrows indicate low tibial mound with sensilla. Black arrows and contour lines show a dentate vestige of the embolickeel. Scale bars: 0.25 mm.