Dichagyris turana cisiliensis, Tang & Zhang & Guo & Luo & Jiang & Pan, 2023

Tang, Jin, Zhang, Gongaote, Guo, Junqi, Luo, Lingxuan, Jiang, Jiamei & Pan, Hongbo, 2023, Contributions To The Taxonomy And Biogeography Of The Genus Dichagyris (Subg. Dichagyris) Lederer, 1867 (Lepidoptera, Noctuidae, Noctuinae) Ii. The Review Of The D. Forficula Species Group, Acta Zoologica Academiae Scientiarum Hungaricae 69 (3), pp. 265-301 : 284-288

Dichagyris turana cisiliensis ssp. n.

http://zoobank.org/ 9B7C903F-77F2-4D9D-A476-1426DDBE96E5 ( Figs 27, 28 , 55–57 , 74 )

Holotype: male, “ Kazakhstan, Chu Ili Mts. , Kopalisai, 600 m, 4.VI.1992, leg. M. Danilevsky ”, slide GYP5785 (coll. PG).

Paratypes. Kazakhstan. 4 males, 1 female, with same data as holotype (coll. PG); 3 males, 2 females, “ Kazakhstan Chu-Ily Mts. | Almaly (sic!) 900 m 5.6.1992 | leg. Danilevsky ”. Slide Nos : VZ9957 , VZ962 , VZ10006 (males), VZ9962 (female) (coll. VZ, RG); 4 males, 4 females, Dzhambul range, Kurdai pass, 1000m, 43°21’N, 75°00’E, 27-28.VI.1993, leg. V. A. Lukhtanov, slide Nos: GYP 5780m, GYP5779f (coll. PG), 1 male, 1 female , Kazakhstan, Kentau , Bayaldir river, 43°37’N, 31°37’E, 586 m, leg. A. Shovkoon (coll. OP). Slide Nos VZ 12001m, VZ11993 f .

Diagnosis. The new subspecies differs externally from the typical specimens of D. turana by the darker fuscous-grey colouration and the more obsolescent pattern. Head, thorax and forewings are rather uniformly dark brownish grey, without ochreous shine. The reniform and orbicular stigmata and the subterminal line are marked with lighter grey scales. The crosslines are marked at the costa with diffuse blackish spots, the cilia are chequered with light grey and blackish scales. The abdomen and the basal part of the hindwings are whitish-grey. The specimens are on average slightly larger than the typical D. turana : 37–38 mm (males), 40 mm (female).

The male genitalia ( Figs 55–57 ) show the taxonomically most important traits. Valvae are longer than those of D. forficula and are only gradually tapering distally, i.e. they have neary parallel margins. Clasper acute, slightly falcate, corona consists of 12–14 setae. Vesica is projecting ventrally, without subbasal diverticulum and cornutus, strongly saccate, not recurved helicoidally.

Female genitalia ( Fig. 74 ). Antrum broad U-shaped, more strongly sclerotised than in the related species; appendix bursae is strongly saccate and much shorter than corpus bursae.

Etymology. The name refers to the region near to the Ili river in Kazakhstan.

Note. A further, slightly darker greyish brown and larger male specimen (41 mm) was found in the Püngeler collection ( MNB) with the labels: “renigera var. turana m v. Tancré 2/1900” (hand written with black ink), “Asia centr. (Issykul) 1894” (printed). It shows the external and genital traits of this subspecies, and it can be geographically allied to this subspecies. Unfortunately, due to the deficient data we desist to consider it as a Paratype of this taxon.

Bionomics and distribution. The new subspecies occurs in semi-desertlike submontane areas near to the border of the Djambul and Almaty regions of Kazakhstan; the moths are on the wing in a relatively early saison (from early June to the end of June).

Dichagyris furiosa furiosa ( Bang-Haas, 1912) , stat. nov.

( Figs 29, 30 , 58 , 59 , 75, 76 )

Agrotis (Lycophotia) renigera var. furiosa Bang-Haas, 1912: 141 .

Type locality: Tadjikistan, Peter The Great Mts , Garm. Lectotype: male, here designated.

Lectotype designation. The male specimen, figured by BANG-HAAS (1922, Taf. XVII: 9) was dissected by Boursin (Slide No. Boursin MB360 ) and is designat- ed here as Lectotype ( Fig. 29 ). The female with the same label data was dissected by Fibiger (slide No. MF2508) and is treated here as Paralectotype ( Fig. 30 ).

Diagnosis. The taxon Agrotis (Lycophotia) renigera var. furiosa Bang-Haas, 1912 was also misidentified until yet, listing as a synonym of D. forficula (see, for instance, http://ftp.funet.fi/ index/Tree_of_life/insecta/lepidoptera/ditrysia/noctuoidea/noctuidae/noctuinae/dichagyris/#8847). According to the short original description the forewings have a violet-reddish colouration with some rosy shade and with prominent pattern. The transverse lines are marked with deep black spots at the costa and the hindwings are blackish grey, with darker marginal area. The underside of wings is nearly evenly blackish grey, the hindwings are sligthly lighter. Wingspan 37–42 mm, females slightly larger.

This description can be completed with the following diagnostic traits: the shape of the forewing is more elongate and acute apically than in the related D. forficula and D. turana , the reddish colouration of the forewings is densely irrorated with blackish-brown scales and the course of the crosslines are more oblique than in D. turana .

Male genitalia ( Figs 58 , 59 ). Valvae are longer and broader than in D. forficula , costa below the cucullus is slightly tapering only, clasper is more elongate and straight than in D. forficula and D. turana . Aedeagus is relatively thick and short, vesica is tubular, not recurved helicoidally, however it has a short triangular subbasal diverticulum armed with a thin and relatively long needle-shaped cornutus.

Female genitalia ( Figs 75, 76 ). The sclerotisation of the antrum is relatively weak and V-shaped. Appendix bursae is much shorter and more saccate than the corpus bursae but less saccate than in D. turana .

Distribution. The nominotypical subspecies of D. furiosa seems to be localised to a relatively restricted area in Tajikistan. The recently collected specimens from “ Tajikistan, Khodshent reg. Mogoltau, 16- 17.05.1996, leg. Lukhtanov” (coll. PG) can also be considered as representatives of the typical subspecies while the specimens from an other mountain system of the Tien- Shan range in Kirghisia belong to a new subspecies, described below.

Figures

Figs 25–32. Dichagyris spp. 25–26 = D. turana turana (Staudinger, 1892): 25 = male, Kirghisia, Issyk-Kul, 26 = female, Uzbekistan, Kyzyl-Kum. 27–28 = D. turana cisiliensis ssp. n., Kazakhstan, Chu-Ili Mts: 27 = holotype, male, 28 = paratype, male. 29–30 = D. furiosa furiosa (Bang-Haas, 1912), Tadjikistan, Peter the Great Mts, Garm: 29 = lectotype, male, 30 = paralectotype, male. 31–32 = D. furiosa griseoerythra ssp. n., Kirghisia, Naryn region, Sarykam-

Figs 53–55. Male genitaliaofDichagyristurana sspp.53–54 =D.turana turana (Staudinger,1892): 53 = Issyk-Kul, coll. Püngeler, slide No.: VZ9551m; 54 = Kirghisia, Osh, slide No.: VZ11987m. 55 = D. turana cisiliensis ssp. n., paratype, Kazakhstan, Chu Ili Mts, slide No.: VZ9751m

Figs 56–58. Male genitalia of Dichagyris spp. 56–57 = D. turana cisiliensis ssp. n., Kazakhstan, Chu Ili Mts: 56 = paratype, slide No.: VZ9957m; 57 = paratype, slide No.: VZ10006m. 58 = D. furiosa furiosa (Bang-Haas, 1912), lectotype, Tadjikistan, Peter the Great Mts, Garm, slide No.: Boursin MB360m

Figs 74–76. Female genitalia of Dichagyris spp. 74 = D. turana cisiliensis ssp. n., paratype, Kazakhstan, Chu Ili Mts, slide No.: VZ9962f. 75–76 = D. furiosa furiosa (Bang-Haas, 1912):

Figs 59–61. Male genitalia of Dichagyris furiosa sspp. 59 = D. furiosa furiosa (Bang-Haas, 1912), Tadjikistan, Alai range, slide No.: VZ11989m. 60–61 = D. furiosa griseoerythra ssp. n.,