Cyana dohertyi eirene Volynkin & N. Singh, 2020

Singh, Navneet, Volynkin, Anton V., Kirti, Jagbir Singh, Datta, Harvinder Singh & Ivanova, Maria S., 2020, A review of the genus Cyana Walker, 1854 from India, with descriptions of five new species and three new subspecies (Lepidoptera: Erebidae: Arctiinae: Lithosiini), Zootaxa 4738 (1), pp. 1-93 : 38

Cyana dohertyi eirene Volynkin & N. Singh , ssp. nov.

( Figs 102, 103 , 205 , 269 )

Type material. Holotype ( Figs 102 , 205 ): ♂, “ Indien, [Uttarakhand] U. P., Kumaon-Himalaya, Bhimtal , 1500 m, 12.IV.1976, leg. F. Smetaсek”, slide MWM 31804 Volynkin (Coll. MWM / ZSM).

Paratypes: INDIA. HIMACHAL PRADESH: 1 ♂, N.W. Himalaya ( Coll. NHMUK) ; 1 ♂, India, H.P., Kan- daghat, 1500m, 9–10.VI.1975, leg. W. Thomas, slide MWM 35885 (♂) Volynkin ( Coll. MWM / ZSM) ; UTTARA- KHAND: 6 ♂, India sept., Kumaon-Himalaya, Distr. NainiTal , BhimTal , 1500 m, leg. F. Smetacek, 10.VII.1979 (1 ♂); 18.IX.1979 (2 ♂); 22.IX.1979 (1 ♂); 24.IX.1979 (2 ♂) ( Coll. MWM / ZSM) ; 2 ♂, 3 ♀, India sept., Kumaon- Himalaya, Distr. NainiTal , BhimTal , 1500 m, Smetacek / Bauer, 31.VIII.1976 (1 ♂, 1 ♀), slide MWM 35881 (♂) Volynkin; 1.IX.1976 (1 ♂, 1 ♀), slide MWM 35880 (♂) Volynkin; 11.IX.1976 (1 ♀) (Coll. MWM / ZSM) ; 3 ♂, 1 ♀, India, Kumaon-Himalaya, Bhimtal , 1500 m, leg. F. Smetaсek 3.V.1975 (1 ♂); 12.VI.1976 (1 ♂), slide MWM 31804 (♂) Volynkim ; 5.IX.1976 (1 ♂); 24.IX.1975 (1 ♀); 1.X.1977 (1 ♂) ( Coll. MWM / ZSM) ; 1 ♂, India , Bhim- tal, 1500 m, 11–20.VI.1975, leg. W. Thomas (Coll. MWM / ZSM) ; 1 ♂, India, Kumaon-Himalaya, Distr. NainiTal , BhimTal , 20–23.V.1981, 1500 m, leg. C. Holzschuh (Coll. MWM / ZSM) ; 2 ♂, 1 ♀, N-India, Bhimtal, 1500 m, 14–30.VII.1979, coll. Dr. Liedgens, slides MWM 31802 (♂), MWM 35884 (♀) Volynkin ( Coll. MWM / ZSM) ; 11 ♂, 2 ♀, India, Kumaon-Himalaya, Distr. NainiTal , BhimTal , 1500 m, leg. de Freina, Himalaya Expedition, 16.V.1971 (1 ♂); 26.V.1971 (2 ♂); 10.V.1971 (4 ♂, 1 ♀); 10.VI.1971 (1 ♂), slide MWM 35883 (♂) Volynkin ; 12.VI.1971 (2 ♂); 18.VI.1971 (1 ♂, 1 ♀) ( Coll. MWM / ZSM) ; 1 ♂, India, Kumaon-Himalaya, Distr. NainiTal , BhimTal , 1500 m, 21.VI.1981, leg. Smetaсek (Coll. MWM / ZSM) ; 3 ♂, Kumaon-Himalaya, Nainital-distr., Bhim- tal, 1500 m, leg. F. Smetaсek, 30.V.1978 (1 ♂), 1.VII.1978 (2 ♂) ( Coll. MWM / ZSM) ; 1 ♂, 5 ♀, Bhimtal, N.- India, 1600 m, 24.VIII.–3.IX.80, Hauenstein, slide MWM 35882 (♀) Volynkin ( Coll. MWM / ZSM) . PAKISTAN. AZAD JAMMU & KASHMIR: 1 ♂, Pakistan, Kashmir, Himalaya Mts. , 20 km В Muzaffarabad , Nara village , 750m, 73°29’E 34°01’N, 20.V.1998, leg. Gy.M. László & G. Ronkay, coll. Museum Witt, slide MWM 31801 (♂) Volynkin ( Coll. MWM / ZSM) ; 8 ♂, 3 ♀, Pakistan, Kashmir, Himalaya Mts. , 30 km N Murree, near Nathia Ghali , Ayubia vil- lage, 2600m, 23.V.1998, leg. Gy.M. László & G. Ronkay, coll. Museum Witt, slides ZSM Arct. 71/2017 ♂, ZSM Arct. 72/2017 ♀ Volynkin ( Coll. MWM / ZSM) .

Etymology. In the Greek mythology, Eirene was one of the Horae, the personification of peace.

Diagnosis. Forewing length is 15–16.5 mm in males and 17–18 mm in females. Cyana dohertyi eirene differs from the nominate subspecies and C. d. mertsana (illustrated by Volynkin et al. (2019)) by its narrower and paler yellow transverse lines. The male and female genitalia of the new subspecies have no significant differences from those of the nominate subspecies. Compared to those of C. d. mertsana (illustrated by Volynkin et al. (2019)), the male genitalia of C. d. eirene have the slightly narrower sacculus with slightly weaker distal process, and the slightly shorter band-like cluster of spinules on the largest diverticulum of vesica. The female genitalia of C. d. eirene differ from those of C. d. mertsana (illustrated by Volynkin et al. (2019)) by the slightly shorter sclerotized posterior section of cospus bursae.

Distribution. Western Himalaya: Pakistan (Azad Jammu & Kashmir) and North India (Himachal Pradesh and Uttarakhand) ( Smetacek 2008, as C. dohertyi ).

Figures

FIGURES 96–103. Cyana spp.: adults. 96, C. gazella, ♂, India, West Bengal (MWM/ZSM); 97, ditto, ♀, India, Uttarakhand (MWM/ZSM); 98, C. dohertyi dohertyi, lectotype ♂, India, Manipur (©NHMUK); 99, ditto, paralectotype ♀, India, Manipur (©NHMUK); 100, ditto, paralectotype ♂, India, Manipur (©NHMUK); 101, ditto, ♂, Nepal (MWM/ZSM); 102, C. dohertyi eirene, holotype ♂, India, Uttarakhand (MWM/ZSM); 103, ditto, paratype ♀, India, Uttarakhand (MWM/ZSM).

FIGURES 203–205. Cyana spp.: male genitalia. 203, C. gazella, India, West Bengal, slide MWM 34492 Volynkin; 204, C. dohertyi dohertyi, Nepal, slide MWM 35886 Volynkin; 205, C. dohertyi eirene, holotype, India, Uttarakhand, slide MWM 31804 Volynkin.

FIGURES 268–272. Cyana spp.: female genitalia. 268, C. dohertyi dohertyi, India, Sikkim, slide NHMUK010313264 Volynkin (©NHMUK); 269, C. dohertyi eirene, paratype, India, Uttarakhand, slide MWM 35882 Volynkin; 270, C. sikkimensis, India, Sikkim, slide MWM 34428 Volynkin; 271, C. divakara, India, Sikkim, slide MWM 34472 Volynkin; 272, C. britomartis, paratype, India, West Bengal, slide MWM 34447 Volynkin.