Tolegnaro kepleri, Álvarez-Padilla & Ubick & Griswold, 2012

Álvarez-Padilla, Fernando, Ubick, Darrell & Griswold, Charles E., 2012, Noideattella and Tolegnaro, Two New Genera of Goblin Spiders from Madagascar, with Comments on the Gamasomorphoid and Silhouettelloid Oonopids (Araneae, Oonopidae), American Museum Novitates 2012 (3745), pp. 1-76 : 69

Tolegnaro kepleri , new species

Figures 342–370 , 377

TYPES: Male holotype and female allotype from Berlese and sifted leaf litter of a tropical dry forest at an elevation of 100 m at Parc National de Namoroka , 17.8 km 329° WNW Vilanandro, 16°22′36″S, 45°19′36″E, Mahajanga, 8–12 December 2002, Fisher-Griswold Arthropod Team, deposited in CASC ( PBI_OON 00036155 ). The species epithet is in honor of the astronomer Johannes Kepler .

DIAGNOSIS: T. kepleri can be differentiated from other Tolegnaro species by the following combination of characters: carapace side microsculpture finely reticulate (figs. 346, 361); male cheliceral paturon inner margin distal portion with long curved modified seta (figs. 348, 363); sternum with radial furrows between coxae I–II, II–III, III–IV, surface finely reticulate, microsculpture only in furrows (figs. 346, 362); leg spines absent (fig. 364).

DESCRIPTION: Male: Total length 1.8. Cephalothorax: Sternum anterior margin with interrupted transverse groove ( fig. 359 ). Mouthparts: Paturon inner margin with recumbent, curved setae (figs. 348, 363). Abdomen: Postepigastric area setae plumose. Legs: Pale orange; tibia I unmodified. Male genital opening slightly protruded, epigastric scutum apodemes large, separated from the anterior book lung opening and not connected by a furrow. Posterior spiracles connected by a groove ( fig. 352 ). Leg spines absent (fig. 364). Tarsi I to IV superior claw teeth not examined in detail. Trichobothria examined with SEM. Tarsal organ with two sensilla visible. Genitalia: Palp cymbium with distal patch of setae. Palp proximal segments pale orange; cymbium pale orange (figs. 347–349). Embolus dorsal lamella and ventral lamella with at least two apophyses each (figs. 367–370). Palp as in figures 365 and 366. Female (PBI_OON 0003325): Total length 1.9. As in male except as noted. Cephalothorax: Mouthparts: Cheliceral paturon inner margin with scattered setae. Legs: Tibia I Emerit’s glands present. Tarsi I to IV superior claw teeth not examined in detail. Genitalia: Genital opening slit shaped ( fig. 353 ). Sperm sac secretory glands openings flat, extending over the receptaculum center. GAp process shaped as a twisted tube ( figs. 354, 355 ). Genitalia as in figures 350 and 351.

MATERIAL EXAMINED: N = 69. Madagascar: Mahajanga: Parc National de Baie de Baly , 12.4 km 337° NNW Soalala, tropical dry forest, 10 m, S16.01000°, E45.26500°, Nov. 26–30, 2002, Fisher-Griswold Arthropod Team, 15 mixed ( CASC PBI _ OON 3044 ) ; 15 mixed ( CASC PBI _ OON 3357 ) ; Parc National de Namoroka , 16.9 km 317° NW Vilanandro, tropical dry forest, 100 m, S16.40666°, E45.31000°, Nov. 12–16, 2002, Fisher-Griswold Arthropod Team, 9 mixed ( CASC PBI _ OON 3325 ) ; 2 mixed ( CASC PBI _ OON 36282) ; Parc National de Namoroka , 17.8 km 329° WNW Vilanandro, tropical dry forest, 100 m, S16.37666°, E45.32666°, Nov. 08–12, 2002, Fisher-Griswold Arthropod Team, 32 mixed ( CASC PBI _ OON 1993 ) . Toamasina: Forêt Clasee Andriantately , rainforest, 530 m, S18.69500°, E48.81333°, Dec. 04–07, 1998, H.J. Ratsirarson, 3 mixed ( CASC PBI _ OON 3838 ) . Toliara: 2.7 km 302° WNW Ste. Luce , littoral forest, 20 m, S24.77166°, E47.17166°, Dec. 09–11, 1998, B.L. Fisher, 1♂ ( CASC PBI _ OON 3889 ) ; Fort Dauphin, Mt. Vasiha , primary forest, 100 m, Feb. 07, 1995, K. Emberton, 1♀ ( MRAC MT. 202. 867 PBI _ OON 8036 ) ; Parc National d’Andohahela , 36.7 km 341° NNW Tolagnaro, montane rainforest, 900 m, S24.76388°, E46.75166°, Jan. 21–25, 2002, Fisher-Griswold Arthropod Team, 2♀ ( CASC PBI _ OON 3871 ) ; Reserve Integrale Naturelle d’Andohahela , parcel 1, 20.0 km SE Andranondambo, 1857 m, S24.56166°, E46.72166°, Nov. 27–Dec. 05, 1995, S. Goodman, 1♀ ( FMNH PBI _ OON 36301) .

Figures

Figs. 342–351. Tolegnaro kepleri, new species. 342. Habitus male, dorsal view. 343. Same, ventral view. 344. Same, lateral view. 345. Cephalothorax, dorsal view. 346. Same, ventral view. 347. Palp male, retrolateral view. 348. Cephalothorax male, anterior view. 349. Palp male, prolateral view. 350. Genitalia female, ventral view. 351. Same, dorsal view. Scale bars = 0.2 mm, except genitalia: 0.1 mm.

Figs. 352–359. Tolegnaro kepleri, new species. 352. Epigastric region male, ventral view. 353. Same female, ventral view. 354. Genitalia female, dorsal view. 355. Same, lateral view. 356. Receptaculum papillae field, anterior view. 357. Pedicel area female, lateral view 358. Endites female, ventral view. 359. Endites male, ventral view. Scale bars = 50 µm, except 356: 10 µm.

Figs. 360–370. Tolegnaro kepleri, new species. 360. Cephalothorax male, dorsal view. 361. Same, lateral view. 362. Sternum male, ventral view. 363. Chelicerae modified setae male, anterior view. 364. Leg 1 female, retrolateral view. 365. Palp male, prolateral view 366. Same, retrolateral view. 367. Embolus, retrolateral view. 368. Embolus, prolateral view. 369. Embolus, ventral view. 370. Embolus, dorsal view. Scale bars = 50 µm except 556–559: 10 µm.