Holmelgonia basalis, JOCQUE & SCHARFF, 1986

Lin, Shou-Wang, Lopardo, Lara & Uhl, Gabriele, 2022, Evolution of nuptial-gift-related male prosomal structures: taxonomic revision and cladistic analysis of the genus Oedothorax (Araneae: Linyphiidae: Erigoninae), Zoological Journal of the Linnean Society 195, pp. 417-584 : 482-484

HOLMELGONIA BASALIS JOCQUÉ & SCHARFF, 1986

( FIGS 19C , 22C , 24C , 36 ; SUPPORTING INFORMATION, FIG. S2A )

Elgonella basalis Jocqué & Scharff, 1986: 12 , figs 18–25 (Dmf).

Elgonia basalis Platnick, 1989: 232 .

Elgonia basalis Scharff, 1990a: 25 , figs 44–46 (mf).

Holmelgonia basalis, Jocqué & Scharff, 2007: 161 .

Examined material: Tanzania, Uzungwa Mts. , Mwanihana Forest above Sanje River, 1700 m, 1♂ 1♀ 15.viii.1982, leg. M, Stoltze & N, Scharff ( ZMUC) ; same locality, 1800 m, pitfall trap, 1♂ 18.viii.1982, leg. M, Stoltze & N, Scharff ( ZMUC) ; same locality, 1800 m, litter, 1♀ 15.viii.1982, leg. M, Stoltze & N, Scharff ( ZMUC) .

Description:

Male ( ZMUC, 15.viii.1982): Total length: 1.72. Prosoma: 0.79 long, 0.61 wide, without modification ( Fig. 19C ). Eyes: AME-AME: 0.02, AME width: 0.04, AME-ALE: 0.02, ALE width: 0.08, ALE-PLE: 0, PLE width: 0.09, PLE-PME: 0.08, PME width: 0.02, PME- PME: 0.03. Clypeus: not hirsute, one sub-AME seta. Sternum: 0.44 long, 0.48 wide. Chelicerae: mastidia absent; stridulatory striae ridged, rows widely and evenly spaced ( Fig. 22C ). Legs: tibia chaetotaxy 2-2-1-1; dorsal proximal macroseta on tibia II, III and IV 2.24, 4.1 and 3.85 times diameter of tibia, respectively; Tm I: 0.60. All metatarsi with trichobothrium. Pedipalp: patella prolateral proximal vertical macrosetae absent; tibia with one prolateral, two retrolateral trichobothria; TPA large, base distant from C base, distal part as wide as base, with two protuberances at tip, each with a seta below ( Fig. 36A, C ); PC base not visible from dorsal view, distal setae close to distal clasp, distal-setae-bearing area wide, distal clasp without striae, clasp extended apically ( Fig. 36A ); T without papillae, PT apical part with long papillae; TS shot, slender ( Fig. 36D ); MSA present; DSA wide, truncated at tip ( Fig. 36A ); EM broad, flat, without papillae, not exceeding ARP; ARP pointed; LER absent; central embolar apophysis present; VRP long, with papillae close to tip; TP long, round at tip; E short and slender, slightly retrolaterally curved ( Fig. 36B ). Opisthosoma: dorsal pattern see Fig. 24C ; PMS with mAP, AC absent; PLS with triad, one AC ( Fig. 36G ).

Female: Total length: 2.04. Prosoma: 0.88 long, 0.62 wide. Eyes: AME-AME: 0.02, AME width: 0.04, AME-ALE: 0.02, ALE width: 0.08, ALE-PLE: 0, PLE width: 0.07, PLE-PME: 0.04, PME width: 0.07, PME- PME: 0.03. Clypeus: not hirsute, one sub-AME seta. Sternum: 0.5 long, 0.51 wide. Legs: tibia chaetotaxy 2-2-1-1; dorsal proximal macroseta on tibia I 2.85 times diameter of tibia, respectively; Tm I: 0.60. Epigyne: Clade 13 characteristic morphology, CO close to dorsal plate posterior margin ( Fig. 36E, F ). Opisthosoma dorsal pattern same as male; PMS with mAP, CY; PLS with triad, one AC, two CY ( Fig. 36H, I ).

Distribution: Tanzania.

Figures

Figure 2. The first part of the three most-parsimonious trees generated from the analysis of Matrix II, with Bremer/ Jackknife support valuesshown beside nodes.The presence/absence of clades in the consensus trees of the most-parsimonious trees of the implied weights analysis with different k values are shown in the box under/on the branches: black for presence when considering all taxa; grey for presence only when Callitrichia convector (Tanasevitch, 2014) and Ca. usitata (Jocqué & Scharff, 1986) are not considered; white for absence.

Figure 19. Male prosomal morphology, lateral. A, ‘Oedothorax’ meghalaya Tanasevitch, 2015 (traced from photograph). B, ‘Oe.’ uncus Tanasevitch, 2015 (traced from photograph). C, Holmelgonia basalis Jocqué & Scharff, 1986 (Jocqué & Scharff, 1986, fig. 18). D, Callitrichia holmi (Wunderlich, 1978) (traced from photograph). E, Ca. picta (Caporiacco, 1949) (Holm 1962, fig. 58). F, Ca. gloriosa (Jocqué, 1984) (traced from photograph). G, Ca. convector Tanasevitch, 2014 (traced from photograph). H, Ca. sellafrontis Scharff, 1990 (traced from photograph). I, Ca. juguma (Scharff, 1990) (Scharff 1990a, fig. 164). J, Ca. uncata (Jocqué & Scharff, 1986) (Jocqué & Scharff, 1986, fig. 130). K, Ca. pilosa (Wunderlich, 1978) (traced from photograph). L, Ca. usitata (Jocqué & Scharff, 1986) (traced from photograph). M, Ca. legrandi (Jocqué, 1985) (Jocqué, 1985, fig. 21). N, Ca. macrophthalma (Locket & Russell-Smith, 1980) (traced from photograph). O, Ca. muscicola (Bosmans, 1988) (traced from photograph). P, Ca. latitibialis (Bosmans, 1988) (traced from photograph). Q, Ca. longiducta (Bosmans, 1988) (traced from photograph). R, Nasoona setifera (Tanasevitch, 1998) (Tanasevitch, 1998a, fig. 3). S, N. crucifera (Thorell, 1895) (Tu & Li, 2004, fig. 6A).

Figure 22. Male chelicerae, stridulatory striae on lateral side. A, ‘Oedothorax’ meghalaya. B, ‘Oe.’ uncus. C, Holmelgonia basalis. D, Callitrichia holmi. E, Ca. picta. F, Ca. gloriosa. G, Ca. convector. H, Ca. sellafrontis. I, Ca. juguma. J, Ca. uncata. K, Ca. pilosa. L, Ca. usitata. M, Ca. legrandi. N, Ca. macrophthalma. O, Ca. muscicola. P, Ca. latitibialis. Q, Ca. longiducta. R. Nasoona setifera. S, N. crucifera. Scale bars 0.05 mm.

Figure 24. Male opisthosoma, dorsal view. A, ‘Oedothorax’ meghalaya Tanasevitch, 2015. B, ‘Oe.’ uncus Tanasevitch, 2015. C, Holmelgonia basalis Jocqué & Scharff, 1986. D, Callitrichia holmi (Wunderlich, 1978). E, Ca. picta (Caporiacco, 1949). F, Ca. gloriosa (Jocqué, 1984). G, Ca. convector Tanasevitch, 2014. H, Ca. sellafrontis Scharff, 1990. I, Ca. juguma (Scharff, 1990). J, Ca. uncata (Jocqué & Scharff, 1986). K, Ca. pilosa (Wunderlich, 1978). L, Ca. usitata (Jocqué & Scharff, 1986). M, Ca. legrandi (Jocqué, 1985). N, Ca. macrophthalma (Locket & Russell-Smith, 1980). O, Ca. muscicola (Bosmans, 1988). P, Ca. latitibialis (Bosmans, 1988). Q, Ca. longiducta (Bosmans, 1988). R, Nasoona setifera (Tanasevitch, 1998). S, N. crucifera (Thorell, 1895). Scale bars 0.1 mm.

Figure 36. Holmelgonia basalis Jocqué & Scharff, 1986. A–D, male right palp, images flipped horizontally. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, F, epigyne. E, ventral view. F, external morphology. G, male posterior median spinnerets and posterior lateral spinnerets. H, female right spinnerets. I, female right posterior lateral spinneret. Scale bars 0.1 mm.