Mitrager noordami, VAN HELSDINGEN, 1985

Lin, Shou-Wang, Lopardo, Lara & Uhl, Gabriele, 2022, Evolution of nuptial-gift-related male prosomal structures: taxonomic revision and cladistic analysis of the genus Oedothorax (Araneae: Linyphiidae: Erigoninae), Zoological Journal of the Linnean Society 195, pp. 417-584 : 485

MITRAGER NOORDAMI VAN HELSDINGEN, 1985

( FIGS 37 , 38O , 39O , 40P; SUPPORTING INFORMATION, FIG. S3C )

Mitrager noordami van Helsdingen, 1985: 353 , figs 1–14 (Dmf).

Examined material: Indonesia: Central Java, Dijeng Plateau, near Gunung Prahu , 2580 m, sifted from litter among mosses, ferns and Ericaceae , 1♂ 1♀ 8.viii.1977, leg. A. Noordam (paratype, Naturalis Museum, Leiden) .

Diagnosis:

Males: This species can be recognized by its unique, prominent prosomal modification ( Fig. 38O ).

Females: Somatic features and coloration similar to other Mitrager and some Callitrichia species, but can be distinguished by the much more convergent borders between the dorsal and ventral plates of the epigyne ( Fig. 37F, G ).

Description:

Male (paratype, Naturalis Museum): Total length: 2.27. Prosoma: 1.13 long, 0.77 wide, with extremely high, ballon-shaped post-PME hump, inter-PME region with a protrution with two round lobes, anterior margin of lobes with row of light-coloured setae, interocular region with two dense, light-coloured setal tufts ( Fig. 38O ). Eyes: AME-AME: 0.03, AME width: 0.05, AME-ALE: 0.06, ALE width: 0.07, ALE-PLE: 0.01, PLE width: 0.08, PLE-PME: 0.1, PME width: 0.07, PME-PME: 0.35. Clypeus hirsute. Sternum: 0.6 long, 0.58 wide. Chelicerae: stridulatory striae imbricated, rows widely and evenly spaced ( Fig. 39O ). Legs: dorsal proximal macroseta on tibia IV 1.80 times diameter of tibia; Tm I: 0.82. Pedipalp: patella prolateral proximal vertical macrosetae absent; tibia with one prolateral, two retrolateral trichobothria; TPS absent; TPA large, conical, distal part narrow and flat, tip curved downwards; PC distal setae close to distal clasp, distalsetae-bearing area slightly wide, distal clasp without striae, clasp extended apically; T without papillae; PT median-long, with small papillae; TS median-long, slender, without papillae; MSA present; DSA wide, round at tip, ventral corner slightly more protruded ( Fig. 37A ); EM broad and flat, retrolateral to LER, with parallel folds extended until anterior margin, exceeding ARP; ARP flat, blunt, short; LER large, bent over E; VRP long, thick at base; R without papillae; TP long,round at tip; E median-long, retrolaterally curved, E tip with prolateral anterior flat extension ( Fig. 37A ). Opisthosoma: dorsal pattern see Fig. 40P ; PMS with mAP, AC absent; PLS with triad, one AC ( Fig. 37H ).

Female (paratype, Naturalis Museum): Total length: 2.48. Prosoma: 1.22 long, 0.84 wide. Eyes: AME-AME: 0.03, AME width: 0.05, AME-ALE: 0.03, ALE width: 0.08, ALE-PLE: 0.01, PLE width: 0.08, PLE-PME: 0.05, PME width: 0.08, PME-PME: 0.12. Clypeus: not hirsute, one sub-AME seta. Sternum: 0.67 long, 0.62 wide. Legs: dorsal proximal macroseta on tibia III 2.39 times diameter of tibia; Tm I: 0.62. Epigyne: Clade 13 characteristic morphology, border between ventral and dorsal plates largely converging toward the middle ( Fig. 37F, G ). Opisthosoma dorsal pattern same as male; PMS one CY, one mAP, AC absent; PLS with triad, two CY, one AC ( Fig. 37I ).

Distribution: Only known from the type locality in Indonesia.

Remarks: The setal tufts at interocular region and inter-PME region are aggregated setae, similar to that found in some Callitrichia species.

Figures

Figure 3. Continuation of Figure 2, showing Clade 36.

Figure 37. Mitrager noordami van Helsdingen, 1985. A–E, male left palp. A, retrolateral view. B, prolateral view. C, dorsal view. D, apical view. E, embolic division, prolateral view. F, G, epigyne. F, ventral view. G, external morphology. H, male posterior median spinnerets and posterior lateral spinnerets. I, female posterior median spinnerets and left posterior lateral spinneret. Scale bars 0.1 mm.

Figure 38. Male prosomal morphology of Clade 55 in Figure 3, lateral.A, Mitrager hirsuta (Wunderlich, 1974) (Wunderlich, 1974, fig. 8). B, M. clypeellum (Tanasevitch, 1998) (traced from photograph). C, M. elongata (Wunderlich, 1974) (Wunderlich, 1974, fig. 13). D, M. cornuta (Tanasevitch, 2015) (Tanasevitch, 2015, fig. 16). E, M. villosa (Tanasevitch, 2015) (Tanasevitch, 2015, fig. 89). F, M. angela (Tanasevitch, 1998) (Tanasevitch, 1998a, fig. 14). G, M. coronata (Tanasevitch, 1998) (Tanasevitch, 1998a, fig. 6). H, M. sexoculorum (Tanasevitch, 1998) (Tanasevitch, 1998a, fig. 19). I, M. globiceps (Thaler, 1987) (traced from photograph). J, M.lineata (Wunderlich, 1974) (traced from photograph).K, M. dismodicoides (Wunderlich, 1974) (traced from photograph). L, M. tholusa (Tanasevitch, 1998) (Tanasevitch, 1998a, fig. 24). M, M. lucida (Wunderlich, 1974) (Wunderlich, 1974, fig. 30). N, M. sexoculata (Wunderlich, 1974) (traced from photograph). O. M. noordami van Helsdingen, 1985 (van Helsdingen, 1985, fig. 1). P, M. unicolor (Wunderlich, 1974) (traced from photograph). Q, M. rustica (Tanasevitch, 2015) (Tanasevitch, 2015, fig. 75). R, M. assueta (Tanasevitch, 1998) (Tanasevitch, 1998a, fig. 1). S, M. malearmata (Tanasevitch, 1998) (Tanasevitch, 1998a, fig. 50). T, M. savigniformis (Tanasevitch, 1998) (Tanasevitch, 1998a, fig. 40). U, M. falcifer (Tanasevitch, 1998) (Tanasevitch, 1998a, fig. 46). V, M. modesta (Tanasevitch, 1998) (Tanasevitch, 1998a, fig. 36). W, M. lopchu (Tanasevitch, 2015) (traced from photograph). X, M. falciferoides (Tanasevitch, 2015) (Tanasevitch, 2015, fig. 35).

Figure 39. Male chelicerae, stridulatory striae on lateral side. A, Mitrager hirsuta. B, M. clypeellum. C, M. elongata. D, M. cornuta. E, M. villosa. F, M. angela. G, M. coronata. H, M. sexoculorum. I, M. globiceps. J, M. lineata. K, M. dismodicoides. L, M. tholusa. M, M. lucida. N, M. sexoculata. O, M. noordami. P,M. unicolor. Q, M. rustica. R, M. assueta. S, M. savigniformis. T, M. falcifer. U, M. modesta. V, M. falciferoides. Scale bars 0.05 mm.

Figure 40. Opisthosoma, dorsal view. A–L, N–Y, male; D, female. A, Mitrager hirsuta (Wunderlich, 1974). B, M. clypeellum (Tanasevitch, 1998). C, M. elongata (Wunderlich, 1974). D, M. cornuta (Tanasevitch, 2015). E, M. villosa (Tanasevitch, 2015). F,M. angela (Tanasevitch, 1998). G, M. coronata (Tanasevitch, 1998). H, M. sexoculorum (Tanasevitch, 1998). I, M. globiceps (Thaler, 1987). J, M. lineata (Wunderlich, 1974). K, M. dismodicoides (Wunderlich, 1974). L, M. tholusa (Tanasevitch, 1998), holotype. M, ‘M. tholusa’, paratype (not conspecific with holotype). N, M. lucida (Wunderlich, 1974). O, M. sexoculata (Wunderlich, 1974). P,. M. noordami van Helsdingen, 1985. Q, M. unicolor (Wunderlich, 1974). R, M. rustica (Tanasevitch, 2015). S, M. assueta (Tanasevitch, 1998). T, M. savigniformis (Tanasevitch, 1998), paratype. U, M. savigniformis, holotype. V, M. falcifer (Tanasevitch, 1998). W, M. modesta (Tanasevitch, 1998). X, M. lopchu (Tanasevitch, 2015). Y, M. falciferoides (Tanasevitch, 2015). Scale bars 0.1 mm.