Clidicus shavrini Jałoszyński, 2019

Jałoszyński, Paweł, Cheng, Zhi-Fei, Brajczewska, Nastazja & Yin, Zi-Wei, 2019, Hidden diversity of giant scydmaenines: five new species and new records of Clidicus in the Philippines (Coleoptera, Staphylinidae, Scydmaeninae), Zootaxa 4658 (3), pp. 571-584 : 580-582

Clidicus shavrini Jałoszyński sp. n.

( Figs 27–33 )

Material studied. HOLOTYPE: ♂: PHILIPPINES, with two labels: “ PHILIPPINES: Mindanao, Davao / prov., Mt. Talomo ( Mts. Apo ), / 1200-1300m, 7°04’40.9’’N, / 125°20’08.3’’E, 27- 28.4.2019, / Shavrin A.V. leg., sifted from wet / litter and debris at banks near / river and in wet litter near / bamboo thickets” [white, printed], “ Clidicus / shavrini Jałoszyński / HOLOTYPUS ” [red, printed] ( MNHW) . PARATYPES (1 ♂, 3 ♀♀): PHILIPPINES : 2 ♀♀, same data as for holotype ; 1 ♂, 1 ♀, “ PHILIPPINES: Mindanao, Davao / prov., Mt. Talomo ( Mts. Apo ), / Catigan , 800-1000m, / 7°01’21.0’’N, 125°22’30.5’’E, / 29.4- 1.5.2019, Shavrin A.V. leg. / Narrow shady valley with stream; / sifted from wet litter at banks near stream / and in dried and wet leaves under bushes” [white, printed] (paratypes in cPJ, cAS) .

Diagnosis. Pronotum with evenly distributed, deep and moderately dense punctures (distances between most punctures subequal to their diameters), lacking median impunctate line and lacking median longitudinal groove, cuticle between punctures glossy; aedeagus with symmetrical parameres, external margin of each broadly and shal- lowly emarginate in subapical region, in lateral view parameres in subapical region abruptly expanded dorsad.

Description. Body of male ( Fig. 27 ) elongate and slender, head flattened, prothorax and elytra strongly convex; legs and antennae long and slender, BL 5.13–5.35 mm; cuticle moderately glossy (more so on head), pigmentation uniformly brown, setae indistinctly lighter than cuticle.

Head ( Fig. 28 ) broadest near posterior third of tempora, HL 0.88–0.90 mm, HW 1.23 mm; tempora in dorsal view slightly more than 3 times as long as eyes, in anterior third straight and slightly divergent posterad, posteriorly strongly and evenly rounded; vertex with a deep subtrapezoidal posteromedian impression and a pair of posterior pits marking sites where dorsal tentorial arms are fused with dorsal wall of head. Sides of vertex convex, frons between antennal insertions nearly flat, entire frons and vertex with a narrow median longitudinal groove, supraantennal tubercles small but distinctly elevated; compound eyes small, nearly round, strongly convex but in dorsal view not projecting from head silhouette, finely faceted. Punctures on sides of vertex and frons fine and sparse, inconspicuous, mesal margin of each eye and posterolateral margin of each supraantennal tubercle with an irregular group of more distinct and denser punctures; supraantennal tubercles and posteromedian vertexal impression virtu- ally impunctate. Setae long, sparse, suberect to erect. Antennae ( Fig. 29 ) slender, scape slightly longer than head, antennomeres II–IX elongate (IX indistinctly so), X–XI about as long as broad; AnL 3.15–3.33 mm.

Labrum with indistinct, shallow and short anteromedian emargination, anterior margin at each side slightly convex, with a pair of barely marked lateral emarginations.

Pronotum with subpentagonal disc and short posterior ‘collar’, broadest near anterior fourth; PL 1.23–1.35 mm, PW 1.23–1.33 mm; ‘collar’ demarcated by a narrow transverse groove with 8–9 small pits; additionally, two larger pits are present on each side of pronotum in its constricted portion; base lacking median longitudinal groove. Punctures on disc distinct, much larger and deeper than those on head, almost evenly distributed, those on central area separated by spaces subequal to diameters of punctures, punctures reducing in diameter and depth toward sides of pronotum. Setae moderately long, sparse, suberect. Cuticle between punctures on pronotal disc as glossy as that on head.

Elytra oval, broadest distinctly in front of middle, EL 3.03–3.10 mm, EW 1.88–2.00 mm, EI 1.55–1.61; each elytron with five dorsal and one lateral rows of large and deep, densely distributed pits, cuticle between pits very finely shagreened, less glossy than that on head, between rows with very fine and sparse setiferous punctures; setae longer and slightly thicker than those on head and pronotum, moderately dense, erect. Apex of each elytron obliquely truncated, with lateral corner not marked, rounded, adsutural corner moderately strongly projecting posterad, sharp-angled and nearly pointed.

Hind wings absent.

Legs long and slender, unmodified.

Aedeagus ( Figs 30–31 ) strongly elongate but moderately slender, AeL 1.40 mm; parameres of equal length and shape, short, in ventral view lateral margin of each broadly and shallowly emarginate in subapical region, apices distinctly divergent distad, rounded, in lateral view parameres in subapical region abruptly expanded dorsad; flagel- lum differentiated from ductus ejaculatorius by a rapid broadening between proximal fourth of median lobe. ‘Sperm pump’ ( Fig. 32 ) 0.68 mm in length, with two large ‘funnels’ of similar shape and diameters.

Female. Externally very similar to male, with slightly shorter antennae; BL 5.03–5.30 mm (mean 5.17 mm); HL 0.88–0.95 mm (mean 0.92 mm), FW 1.25–1.35 mm (mean 1.32 mm), AnL 3.00– 3.13 mm (mean 3.06 mm); PL 1.18–1.30 mm (mean 1.24 mm), PW 1.23–1.33 mm (mean 1.28 mm); EL 2.98–3.05 mm (mean 3.01 mm), EW 1.83–2.05 mm (mean 1.92 mm), EI 1.49–1.63.

Spermatheca ( Fig. 33 ) with ovoid capsular part, short subconical projection connected to ductus spermathecae, width of capsular part 0.15 mm; accessory gland with connecting duct conspicuously long; ductus spermathecae strongly entangled.

Distribution. Central-southern Mindanao.

Etymology. This species is dedicated to Alexey Shavrin, a specialist on omaliine staphylinids and a collector of the type series of this interesting species.

Remarks. Clidicus shavrini is most similar to C. occisor ; differences were given in remarks for the latter spe- cies. Adults of this species were collected by sifting leaf litter and other debris near a river ( Figs 35–36 ), but also in a more disturbed habitat of a banana plantation from under heaps of drying grasses and wet leaves ( Fig. 37 ). The latter collecting circumstances seem rather unusual for Clidicus .

Figures

FIGURES 27–33. Clidicus shavrini Jałoszyński sp. n. Habitus of male (27); head of male in dorsal view (28), right antenna of male in dorsal view (29); aedeagus in ventral (30) and lateral (31) views; ‘sperm pump’ (32); spermatheca (33).

FIGURES 34–37. Collecting sites of Clidicus forceps Cheng et al. sp. n. (34) and C. shavrini Jałoszyński sp. n. (35–37). Can- dalaga Mts., Compostela Valley (34); Mt. Talomo (35–37).