Calapnita magaseng, Bernhard A. Huber, 2017

Bernhard A. Huber, 2017, Revision and cladistic analysis of the Southeast Asian leaf-dwelling spider genus Calapnita Simon (Araneae, Pholcidae), Zootaxa 4219 (1), pp. 1-63 : 37

Calapnita magaseng sp. nov. Figs 122–123 , 156–163

“ Calapnita vermiformis ” (misidentification; only specimens from Semengoh Arboretum): Huber 2011: 48.

Diagnosis. Easily distinguished from other species of vermiformis group by strong lateral processes on male chelicerae ( Fig. 160 ); also by tip of procursus (ventral flap strongly protruding towards ventral; Figs 156–157 ). From C. bario and C. bariengi also by distal cheliceral apophyses clearly bipartite ( Fig. 160 ); from several species also by narrow and distinct epigynal ‘knob’ ( Fig. 162 ), and by oval pore plates very close together ( Fig. 163 ).

Etymology. The species name is derived from the type locality; noun in apposition.

Material examined. Holotype. MALAYSIA-BORNEO: ♂, ZFMK ( Ar 16026), Sarawak, Mount Penrissen near Borneo Highland Resort, Ma Gaseng Trail (1.124– 1.127°N, 110.217°E), 870–930 m a.s.l., underside of leaf, 14.vii.2014 (B.A. Huber, S.B. Huber).

Other material. MALAYSIA-BORNEO: 2♂ 4♀, ZFMK (Ar 16027), same data as holotype ; 5♀ in absolute ethanol, ZFMK (Bor 219), same data . 1♂ 1♀, ZFMK ( Ar 16028), Sarawak, Semengoh Arboretum, Masing Trail (1.397– 1.399°N, 110.317– 110.322°E), 60–80 m a.s.l., undersides of leaves, 17.vii.2014 (B.A. Huber).

Description. Male (holotype)

MEASUREMENTS. Total body length 4.6, carapace width 0.7. Leg 1: 29.1 (7.2 + 0.3 + 6.5 + 12.6 + 2.5), tibia 2: 5.1, tibia 3: 3.2, tibia 4: 5.6; tibia 1 L/d: 108. Distance PME-PME 200 µm, diameter PME 90 µm, distance PME- ALE ~30 µm; no trace of AME.

COLOR. Entire animal pale gray to whitish, legs slightly yellowish with brown patellae and tibia-metatarsus joints.

BODY. Habitus as in Fig. 122 ; ocular area barely elevated, each triad on very low hump; carapace without median furrow; clypeus unmodified; sternum slightly wider than long (0.50/0.44), unmodified.

CHELICERAE. As in Figs 160–161 , with bipartite apophyses near lamellae (proximal processes more distinct in holotype than in drawn male), pair of strong lateral processes; without modified hairs; without stridulatory ridges.

PALPS. In general similar to C. vermiformis (cf. figs 139 and 140 in Huber 2011); trochanter apophysis as in Fig. 159 ; femur as in Fig. 158 , ventral processes widely spaced (distal process at 51% of femur length); tibia length/diameter 0.60/0.28; procursus as in Figs 156–157 , with simple ventro-distal flat sclerite; bulb length 0.39; embolus length 0.70; embolus tip with transparent fringes.

LEGS. Without spines and curved hairs; few vertical hairs; retrolateral trichobothrium on tibia 1 at 2%; prolateral trichobothrium absent on tibia 1, present on other tibiae; tarsus 1 pseudosegments barely visible in dissecting microscope.

Male (variation). Tibia 1 in 3 other males: 6.7, 6.8, 7.0. Proximal processes of bipartite cheliceral apophyses variably long.

Female. In general similar to male; eye triads slightly closer together (distance PME-PME 180 µm). Tibia 1 in 5 females: 5.5–6.2 (mean 6.0). Epigynum as in Fig. 162 , weakly sclerotized triangular plate with small and slender anterior ‘knob’; internal genitalia as in Fig. 163 , with pore plates close together.

Distribution. Known from two localities in western Sarawak only ( Fig. 283 ).

Figures

FIGURE 283. Known distribution of the Calapnita vermiformis group, part 1 (cf. Fig. 284). The Java record of C. lawangan is based on specimens assigned tentatively.

FIGURES 120 – 127. Live specimens, Calapnita vermiformis group. 120 – 121. C. bario sp. nov., ♂ and ♀ with egg-sac from Bario. 122 – 123. C. magaseng sp. nov., ♂ and ♀ with egg-sac from Mount Penrissen. 124 – 125. C. saluang Huber, ♂ from Berastagi and ♀ with egg-sac from Phanom Bencha. 126 – 127. C. vermiformis Simon, ♂♂ from Mount Isarog and Mount Banahaw.

FIGURES 156 – 171. Calapnita magaseng sp. nov. (156 – 163; ZFMK, Ar 16027) and C. dayak sp. nov. (164 – 171; ♂: RMNH, ARA 17421; ♀: RMNH, ARA 17424, 17427). 156 – 157, 164 – 165. Left procursi, prolateral and retrolateral views. 158, 166. Left palpal femora, prolateral views. 159, 167. Left palpal trochanters, retrolateral views. 160 – 161, 168 – 169. Right male chelicerae, frontal and lateral views. 162, 170. Epigyna, ventral views. 163, 171. Cleared female genitalia, dorsal views. Scale lines: 0.3 mm.