Physodeutera, Lacordaire, 1842

Moravec, Jiří & Trýzna, Miloš, 2021, New or rare Madagascar tiger beetles- 23. Physodeutera (Toxoma) lokobensis sp nov., a new species close to Ph. (T.) conturbata Moravec, Ph. (T.) sulcoprothoracica (W. Horn) and Ph. (T.) dubia (Mařan), with revised type designation of the latter (Coleoptera: Cicindelidae), Zootaxa 5060 (2), pp. 151-182 : 154-155

Key to species of Physodeutera ,

subgenus Toxoma

(individual females may be barely identifiable in some species)

1 Apex of antennomere 4 dilated in male. Elytra entirely immaculate in both sexes. Body medium-sized, bright metallic blue with green, greenish-bronze and violet reflections. Aedeagus long with elongate basal half, apical portion ventrally arcuate-emarginated, forming ventrad-directed, rounded apex............................................................... 2

- Antennomere 4 normally shaped. Body medium-sized to large. Elytra with at least humeral macula in male.Aedeagus variously shaped.............................................................................................. 3

2 Antennomere 4 (in male) simply clavate-dilated. Lateral margins of pronotal disc almost parallel, slightly emarginated in middle. Pronotum slightly more elongate with rather rounded, less parallel lateral margins of disc. Elytra bright green-blue. Aedeagus moderately enlarged in middle.............................. Ph. ( T.) lobicornis lobicornis Moravec, 2000

- Antennomere 4 (in male) more distinctly dilated, eccentric-spatulate. Elytra deep green-blue. Aedeagus more distinctly enlarged in middle.................................................. Ph. ( T.) lobicornis nosybensis Moravec, 2000

3 Elytra with conspicuous or well noticeable velvety black shadowy zone on elytral disc; punctures rather coarse and occasionally anastomosing on anterior elytral half, much finer or almost effaced on posterior elytral half...................... 4

- Elytra without noticeable velvety black zone on elytral disc (yet inconspicuous shadowy areas may diffuse on elytral disc or various elytral areas)................................................................................... 5

4 Body medium sized. Elytral disc with conspicuous, well-delimited velvety black shadowy zone in middle, remaining surface metallic black with feeble or bright, blue, green-blue, or bronze reflections, or with chatoyant, gold-bronze, or purple lustre on posterior declivity; whitish maculation consisting of humeral-lateral lunule (absent in female) and sublateral-median macula. Antennomere 4 in male ochre testaceous with black-brown base and apical third. Aedeagus short, attenuated into slightly ventrad-bent, rather wide, rounded apex...................................... Ph. ( T.) subtilevelutina ( W. Horn, 1934)

- Body larger. Elytral disc with faint, yet notable velvety-black shadowy zone diffusing from central area towards outer elytral margins; remaining surface deep cyaneous-blue and green-blue with strong violaceous lustre laterally, sometimes with indistinct bronze lustre; only whitish humeral macula in male, which is in female only indicated as a small, dark, and almost invisible spot, or entirely absent. Antennomere 4 black with basal half light brownish or dark brown. Aedeagus with notably short apical portion towards very short, regularly rounded apex........................ Ph. ( T.) sulcoprothoracica (W. Horn, 1913)

5 Elytra with only white humeral macula in male, female elytra immaculate or with only inconspicuous, darkened humeral macula. Aedeagus rather stout (including its basal portion).................................................... 6

- Elytra also with white sublateral-median macula, punctate throughout, punctures coarser and anastomosing on anterior elytral half, much finer on posterior half. Aedeagus with narrowly elongate basal half..................................... 8

6 Antennomere 4 in male with two basal thirds ivory to ochre-testaceous, elytra bright green-blue, only lateral areas and apices violaceous........................................................................................... 7

- Antennomere 4 entirely black or black-brown in both sexes. Elytra rather coarsely punctate on their anterior half, posterior half with finer and notably spaced punctures yet not entirely effaced, deep cyaneous-violaceous with brighter greenish and indistinct bronze lustre within discal impression; elytral disc with only barely noticeable, feebly diffusing shade. Pronotal disc distinctly attenuated anteriad in male. Aedeagus apex short, regularly rounded.................. Ph. ( T.) conturbata Moravec, 2002

7 Elytra distinctly, almost equally punctate throughout, lacking any shadowy area on elytral disc. Pronotal disc less notably attenuated anteriad in male. Aedeagus apex regularly rounded yet narrower. Female unknown.... Ph. ( T.) dubia ( Mařan, 1942)

- Elytral punctation finer, particularly so and spaced on posterior half of elytral disc, sometimes with indistinct shade in middle. Aedeagus apex wider, rounded but almost triangular-topped (ventrally moderately arcuate and dorsally obliquely sloped towards indistinct emargination)..................................................... Ph. ( T.) lokobensis sp. nov.

8(5-) Pronotum elongate. Elytra predominantly metallic green to green-blue, punctures coarser on anterior elytral half; whitish humeral macula combined with subhumeral-lateral lunule in male, while in female only subhumeral lunule present. Aedeagus with notably narrowly elongate basal half, dorsally enlarged apical half, while ventral margin straight; internal sac notably simple.................................................................... Ph. ( T.) sikorai (W. Horn, 1896)

- Pronotum approximately as long as wide with notably wider disc. Elytra predominantly deep cyaneous, distinctly punctate throughout; whitish humeral macula in male simple, absent in female, yet obscurely indicated subhumeral appendage may occur in both sexes; lateromedian macula smaller. Aedeagus shorter, its apical half more enlarged and apex slightly directed ventrad; internal sac more armed, with distinct, longitudinal-sigmoid ventral piece with clavate apex......................................................................................... Ph. ( T.) breviformis (W. Horn, 1904)