Omophron (Omophron) capense, Gory, 1833

Anichtchenko, Alexander & Valainis, Uldis, 2023, Revision of the subgenus Omophron (s. str.) Latreille, 1802 (Coleoptera: Carabidae Omophron) of the Afrotropical region, Zootaxa 5284 (2), pp. 201-246 : 218-222

Omophron (Omophron) capense s. str.

Loc. Typ.: Cap de Bonne Espérance.

= Omophron suturale Guérin-Méneville, 1842: 24 Loc. Typ : Cap de Bonne Esperance.

= Omophron dominicense Chaudoir, 1868: 58 Loc. Typ. : “S Domingo” [label error]. Syn. n.

Material examined. BOTSWANA: Botswana 1922 (1 ex, ETHZ) ; Bechuanaland , 1922 (1 ♁, BMNH) ; 20 km N Maun , 16.01.1978, Holm leg. [-20.009395, 23.416671] (1 ♀, TMSA) . MOZAMBIQUE: Beira , A. Bodong leg. [- 19.769238, 34.808530] (1 ♁, ETHZ) . NAMIBIA: Windhoek, Eros Mt. , 1600 m, 22°34’S–17°06’E, 10.09.1974, Endrody & Younga leg. [-22.566270, 17.102200] (1 ♁, TMSA) ; Oshakati, 12.1982, Holm & Scholtz leg. [- 17.777564, 15.707188] (1 ♁, TMSA) ; Tsumkwe, 1151 m, 19°35’S–20°29’E, 10.03.2006, Muller leg. [-19.580891, 20.485568] (1 ♀, TMSA) ; S. Africa (W52), Swakop R. , 3 mls. S. Okahandja , 07.04.1972 [-22.045604, 16.909570] (2 ♁, 3 ♀, BMNH) ; Mariental, Hardap dam, 10– 14.04.1972, Strodym & Jones leg. [-24.498859, 17.861124] (1 ♁, TMSA) ; Fish River mouth, shore washing, 28°05’32’’S,17°10’19’’E 01.09.2002, Uhlig leg.[-28.096683, 17.172637] (1 ♀, 1 ♁, ZMHB) ; Namibia-exp. ZMB 1992 , Windhoek: Daan Viljoen sparlich bewachsenes Ufer , 22°26’S / 16°53E, 13–15.ii.92, leg. M. Uhlig [-22.433373, 16.883290] (3 ♁, 1 ♀, ZMHB) ; Voigts Farm, Okaparakaha , 90km NO von Windhoek, 2–4.iii.2001, lux, leg. U. Gröllner [-22.202034, 17.407282] (1 ♀, ZMHB) ; Khomas Str. zw. Aris u.Brack Auas Game Lodge D 1463, 1780m S22°47’46.9 E17°19’45.4, leg. A. Puchner 5.III.2016 [-22.796381, 17.329149] (1 ♁, 7 ♀ PSC) ; Otjozondjupa zw. Osire u. Otjiwarongo Osire , Burgkellefarm , S21°06’42.2 E17°26’21.1, 1360m, 27.II.2018, leg. A. Puchner [-21.111231, 17.440360] (1 ♁, PSC) ; Maltahöhe 12.X.2004, Werner [- 24.850377, 16.953813] (1 ♁, PSC) ; Zoutpansberg, 13.11.2000, Werner leg. [-23.000817, 29.780251] (3 ♀, PSC) ; 29km S Ondangwa Nakambale Museum , Oshikoto prov., 17°59.024S, 16°01.358E, 25.XI.2012, P. Schüle leg. [- 17.981951, 16.024006] (1 ♀, PSC) . REPUBLIC OF SOUTH AFRICA: Ex Musaeo Chaudoir , Type capensis Gory [red label], ♀ [blue square] [-34.355952, 18.479917] (Holotype ♀, MNHN) ; Ezemvelo Nat. Res., 1374 m, 25°42’S– 29°01’E, 27.01.2005 [-25.700396, 29.017268] (2 ♀, TMSA) ; Ezemvelo NR, 20 km NE Bronkhorstspruit , 1190m, 25°42’S–29 °00’E, 19– 20.01.2008, P. Schüle leg. [-25.705101, 28.991184] (6 ♀, 2 ♁, PSC) ; Transvaal Klaserie, 23.01.1999, P. Schüle leg. [-24.523440, 31.064283] (1 ♀, 1 ♁, PSC) ; Badplaas, 1000m, 01.12.2000, P. Schüle leg. (1 ♁, 1 ♀, PSC) ; Vaal River, 1250 m W of Bothaville , 22.12.2007, M. Snizeg. leg [-27.386549, 26.515146] (1 ♀, DUBC; 7 ♀, 2 ♁, PSC) ; W of Bothaville, Vaalriver , 26.X.2009, M. Snizeg. leg (11 ♁, 5 ♀, PSC) ; Lindani Nature Res., nr. Vaivaater , 1190m, 04– 06.01.2008, P. Schüle leg. (1 ♀, PSC) ; KwaZulu-Natal, 35km N Port Edward , 26.12.2009 A.Kudrna leg, (2 ♀, PSC) ; M’fongosi, Zululand, Misses Jones , Feb. 1923 [-28.703599, 30.834583] (1 ♁, SAM-COL-A006504 ; 2 ♁, 3 ♀, SAM-COL-A08085 ) ; idem, but Dec. 1922 (1 ♁, SAM-COL-A006505 ) ; idem, but Dec.– Feb. 1935 (1 ♀, SAM-COL-A018086 ); Natal, Malvern ., C. N. Barker 1909 (2 ♁, SAM-COL-A006501 ); Natal, Estcourt , Haviland , 1894 (1 ♀, SAM-COL-A006506 ); R. S. Africa, 14.xi.1993, 33°24’S / 22°40’E, Cape Province, Grootswartberg ca. 10km SE Swartbergpass , leg. M Uhlig (1 ♀, ZMHB) ; South Africa, KwaZulu-Natal, Cathedral Peak NR, Didima (camps), 28°57.4’S 29°14.4E 1420m, 19–20.XI.2006, P. Bulirsch ( PBC) ; S. Africa Rep., WC, Biedouw valley, Enjo Nature Farm , 32°08’S 19°17’E, 285m, 28– 29.08.2017 [-32.140490, 19.287212] (1 ♁, DUBC) ; Wiedouw farm, 31°43’S–18°43’E, 19.08.1983, Endrody & Penriht leg. [-31.714827, 18.706439] (1 ♀, 1 ♁, TMSA) ; Soutpans, Tswaing , 25°24’S–27°33’E, 04.12.1996, Muller leg. [-25.395941, 27.567688] (2 ♀, TMSA) ; Groot Toren farm, 31°20’S–19°44’E, 15.09.1994, Endroy & Younga leg. [-31.336277, 19.741493] (5 ♁, 1 ♀; TMSA) ; Nuweberg, 34°00’S–19°06’E, 13.11.1973, Endrody & Younga leg. [-34.001429, 19.100473] (1 ♀, TMSA) ; Villiersdorp, 20.11.1968 [-33.979608, 19.278715] (1 ♁, 1 ♀, TMSA) ; Le Cap (4 ♁, MRAC) ; Donker hoek, 13.03.1980, A.P. du Toit leg. [-25.799967, 28.459624] (1 ♁, TMSA) ; Beaufort Distr., Tldburg Toll , berges vaseuses, Kat. Riv ., 01.1961, Leleup leg. (1 ♀, MRAC) ; Beaufort Distr., Katberg , bancs sabl. Ruisseau , 01.1961, Laleup leg. [-32.524080, 26.701004] (11 ♁, 7 ♀,); Sasolburg, 18.10.1996, Kruger leg. (5 ♁, 3 ♀, TMSA) ; Maghaleen 03– 07.03.1969, Jones & Snyman leg. (3 ♁, 5 ♀, TMSA) ; Oranjekrag, 26– 30.10.1970, Snyman & Jenes leg. (1 ♁, TMSA) ; Ladybrand, Italy farm, 09.– 10.11.1970, Snyman & Jones leg. (1 ♁, 1 ♀, TMSA) ; Ladybrand, 12.11.1970, Snyman & Jones leg. (1 ♀, TMSA) ; Transvaal Rodeplaat Dam, 25°37S–28°23E, 14.08.1974, Strydom leg (1 ♁, TMSA) ; Transvaal Rodeplaat, 17.– 18.03.1960, Neubecker leg (1 ♀, TMSA) ; idem but 08– 10.10.1960 (1 ♁, TMSA) ; idem but 30.10.– 10.11.1960 (2 ♀, TMSA) ; Cullinan Premier Mine Game Res., 25°40’S–28°29’E, 17.01.2002 (1 ♀, TMSA) ; Ezemvelo Nat. Res., 1350 m, 25°42’S–29°00’E, 19.01.2008, Muller leg. (1 ♁, 1 ♀, TMSA) ; Pretoria, Roodeplaat , 17– 18.10.1960, Neubecker leg. (1 ♀, TMSA) ; Bundu Inn, 25°28’S–28°55’E, 24.03.1974, Endrody & Younga leg. (4 ♀, TMSA) ; Rivonia, 03.11.1967, Russel leg. (1 ♀); Eastern Cape Province (1 ♀, ETHZ) ; Doornkraal, 33°39’S–24°22’E, 04.12.1995, Bellamy leg. (1 ♁, TMSA) ; Elizabeth port, 1917, Marshall leg. (1 ♀, BMNH) ; Port Elizabeth (10 ♁, 8 ♀, ETHZ; 1 ♀, ZIN); Port Elizabeth, Cape Colony , Bänninger leg. (1 ♁, MRAC) ; Pt. Elizabeth, Kapland (1 ♀, MNHN) ; Elizabeth Port, Brauns leg. (6 ♀, 2 ♁, ZMHB) ; Amatole, Isidenge For. St. , 32°41’S– 27°16’E, 22.11.1987, Endrody & Younga leg. (1 ♁, TMSA) ; Little Karroo Baviaans kloof, 33°38’S–24°15’E, 27.10.1993, Endrody & Younga leg. (4 ♁, 2 ♀, TMSA) ; Middelberg, 12.1961 (3 ♁, 2 ♀, TMSA) ; Transkei Port St Jones, Silaka , 31°33’S–29°30’E, 23.11.1987, Endrody & Younga leg.(1 ♀, TMSA) ; Transkei, The Haven , 24.02.1985, Duke leg. (1 ex, TMSA) ; Transkei, Longeni Forest , 13.12.1984, Duke leg. (1 ♀, TMSA) ; Uitenhage, 1904, Marshall leg. (1 ♁, BMNH) ; 1917, O’Neil leg. (1 ♀, BMNH) ; Katberg , 4000 ft., 14.– 26.11.1932, Turner leg. (1 ♀, BMNH) ; Natal, Reiswnik leg (1 ♀, SMTD) ; Umfolosi Game Reserve , 02.1967, J. & L. Lauw-Bourguin (1 ♁, 2 ♀, TMSA) ; Natal Midlands, Karkloof , 29°18’S–30°13’E, 02.12.1989, Endrody & Klimaszew leg. [-29.300300, 30.217667] (1 ♀, TMSA) ; idem but 03.12.1989 (1 ♁, TMSA) ; idem but 13.12.1989 (1 ♁, TMSA) ; Estcourt, 1917, Marshall leg. (1 ♁, BMNH) ; Weenen, 01.1924, Thomasset leg. (1 ♁, BMNH) ; Weenen, 09.1929, Thomasset leg. (1 ♀, MRAC) ; Natal: Drakensberg: Dragons Peak Park 29°02’S / 29°26E, 8.ii.1994, leg. M. Uhlig (1 ♀, ZMHB) ; Natal: Itala Game Reserve , 27– 29.1.1994, 27°30’S, 31°20’E, lux, U. Göllner leg. (1 ♁, ZMHB) ; Cape Province, Swellendam , Bontebok Nat.P. 2– 3.1.1993, leg. F. Koch (1 ♁, ZMHB) ; Royal NP, Tugela River banks, 28°43’S, 28°53’E, sievings (2 ♁, 5 ♀); Penge , 13– 17.11.1972, Strydom leg. (2 ♁, 3 ♀, TMSA) ; Rhenosterpoort , 25°45’S–28°55’E, 19.02.1975, Pot & Scoble leg. [-25.753020, 28.957033] (1 ♁, TMSA) ; 01.04.1986, Schuze leg. (1 ♀, TMSA) ; Waterberg, Farm 223, 24°11’S–27°50’E, 11.02.1976, Strydom leg. (4 ♁, 2 ♀, TMSA) ; Waterberg , Matlabas river, 24°25’S–27°36’E, 06.10.1995, Endrody & Bellamy leg. (1 ♁, 2 ♀, TMSA) ; Waterberg, Geelhoutbush farm, 24°22’S–27°33’E, 19.12.1995, Bellamy leg. (7 ♁, 9 ♀, TMSA) ; Waterberg, Matlabas river, Hippo P., 24°21’S–27°38’E, 07.10.1995, Endrody & Bellamy leg. (6 ♁, 5 ♀, TMSA) ; Ocoloco , 05.– 13.12.1958, G. Van Son leg. (2 ♁, TMSA) ; Soutpansberg , 23°00’S–29°47’E, 14.03.1973, Endrody & Younga leg. (1 ♀, TMSA) ; Letaba , 05.10.1980, Honiball leg. (1 ♁, 1 ♀, TMSA) ; N. Tvl.: Letaba Valley , Tzanen Dist., 12.1958, Capener leg. (1 ♁, MRAC) ; Transvaal, Letaba Distr. , Letaba Riv., berges, 09.1960, Leleup leg. (2 ♁, 2 ♀, MRAC) ; Thabaphaswa , 24°00’S–28°55’E, 14.12.2003, Muller leg. (2 ♁, 2 ♀, TMSA) ; Nylsvley , 07.10.1977, Scholtz leg. (2 ♁, 1 ♀, TMSA) ; N. Transvaal, Nylsvley, Smith farm, 24°40’S–28°42’E, 20.07.1975, Endrody-Younga leg. (2 ♁) ; idem but 27.08.1975 (1 ♀) ; idem but 17.09.1975 (2 ♁) ; idem but 13.12.1975 (2 ♁, MRAC) ; N. Tvl.: Gravelotte Letsitele Valley , 16– 12.12.1958, Capener leg. (1 ♁, 2 ♀, MRAC) ; Elandshoek, 09.1947, Capener leg. [-25.500568, 30.699013] (1 ♀, TMSA) ; Nelshoogte, galery for below St. , 25°51’S–30°53’E, 04.12.1986, Endrody & Younga [-25.849900, 30.885138] (1 ♁, 1 ♀, TMSA) ; Kopees Kraal, Lydenberg , 25°07’S–30°12’E, 05.12.1996, Endrody & Younga leg. [-25.114782, 30.211891] (4 ♁, TMSA) ; idem but 06.12.1996, (3 ♁, TMSA) ; Helena Farm , 1091 m, 25°02’S–30°07’E, 22.10.2001, TMSA staf leg. [-25.033134, 30.118736] (1 ♁, TMSA) ; idem but 11.02.2002 (1 ♁, TMSA) ; Barberton, Rendall leg. (1 ♁, BMNH) ; Transvaal, Barberton Distr. , Suid Kaap Riv., rives sabl. argil., 10.1961, Leleup leg. [-25.608914, 31.238459] (7 ♁, 10 ♀, MRAC) ; Transvaal, Lydenburg Disr., Ohrigstad , 1200m, 21.03.1962, Leleup leg. [-24.749446, 30.543391] (4 ♁, 6 ♀, MRAC) ; Botshabelo, Transvaal [-23.434354, 30.529504] (1 ♁, MNHN) ; RSA, Northern Prov., Soutpansberg Mts., Medike res., 4– 6.12.2003 R. Kmeco leg [-22.993408, 29.611937] (3 ♁, 3 ♀, DUBC, RKC) ; Warrenton , 09.03.1978, F. De Moor leg. [-28.110515, 24.843204] (1 ♁, TMSA) ; Upington, Schüle leg. [-28.473178, 21.230935] (3 ♁, 5 ♀, SMNS) ; Nietverdiend , 08.01.1978, Holm leg. [-25.035948, 26.154621] (1 ♀, TMSA) ; Ingwavuma River , valee rives, 10.1961, Leleup leg. [-27.078750, 32.016142] (2 ♁, 8 ♀, MRAC) . O. capense f. suturalis: Dunbrody, Esper leg. (1 ♀, ISNB) ; Dunbrody (9 ♁, 7 ♀, ISNB) ; Dunbrody (4 ♁, 3 ♀, MRAC) ; Dunbrody [-33.474672, 25.552932] (1 ♁, SAM) ; Dunbrody pres Grahamstown, Cap de B. Esper . [-33.328017, 26.563948] (1 ♁, ISNB) ; Algoa Bay, Capland, Brauns leg. 1.12.92 [-33.849003, 25.592648] (1 ♀, TMSA; 1 ♁, 4 ♀, SAMC; 1 ♀, SMTD); Cap. , Gory leg. (1 ♀, 2 ♁, MNHN) , Cap. B. Esp . (1 ♀, MNHN) ; Cap de Bonne Esperance (1 ♀, MRAC) ; Le Cap, Burgeon leg. (1 ♀, MRAC) ; Cape Town (1 ♁, 1 ♀, SMTD) ; Cape Town, Table Bay (1 ♁, ZIN) ; Cubha Lake , 11.1980, Oberprieler leg. [-28.848655, 31.943981] (1 ♁, TMSA) ; Mtunzini, 11.1980, Oberprieler leg. [-28.948662, 31.775811] (1 ♁, TMSA) ; Durban, Tongaat beach, 29°35S–31°11 E, 18– 30.01.1992, Endrody & Younga leg. [- 29.578230, 31.181660] (1 ♁, TMSA) ; Durban, 01.1939, Kobrow leg. [-29.884271, 31.019884] (1 ♁, TMSA) ; Durban, 1903, Marley leg. (1 ♁, 1 ♀, BMNH) ; Durban, 1917, Marshall leg. (1 ♀, BMNH); Zululand, Mtubatuba , 28°22’S–32°19’E, 04.04.1974, Endrody — Younga leg. [-28.367091, 32.318099] (15 ♁, 11 ♀, TMSA) ; Zululand, Empangeni , 23.01.1975, Ereavel leg. [-28.768451, 31.885523] (1 ♁, TMSA) ; “S Domingo” [label error], “Ex Musaeo Chaudoir”, “ Omophron dominicense Chaud. Holotype, det. George E. Ball ”, “Type” [red square], Omophron dominicense Chaudoir, 1868 = O. suturalis Guerin-Meneville 1829 det. S.W. Nichols 1983” (♁, MNHN). ZAMBIA: 30 km NW Sesheke , S17°17´21 E24°09´08, 13.01.2010, P. Schüle leg. [-17.289167, 24.152222] (1 ♁, 1♀ PSC); N. Rhodesia, Namwala, Dollman leg. [-15.999410, 26.138941] (8 ♁, 5 ♀, BMNH). ZIMBABWE: Salisbury, Mashonaland, 1908, Marshall leg. [-17.927256, 30.875280] (3 ♁, BMNH) ; Harare, Bodong leg. (5 ex, ETHZ) ; Mashonaland (1 ♁, MNHN) ; S. Rhodesia, Marandellas , 02.1962, 18°10’S, 31°36’E, Weir leg. [-18.167591, 31.599725] (2 ♁, 3 ♀, BMNH) ; S. Rhodesia, Khami (Bulawayo) (a la lumiere) 10.01.1954, Cooke leg. [-20.151822, 28.428814] (1 ♁, 2 ♀, MRAC) ; „ Rhodesia, Sebakwe, Dods 190”, „O. picturatus”, „ var oligoxanthus ”, SAM-COL-A018083 , [-18.862753, 29.836817] (3 ex, SAMC) ; Rhodesia, Sebakwe, Dods 1902, SAM-COL-A052163 (3 ♁, 1 ♀, SAMC) .

Differential diagnosis. Similar to O. picturatum and O. oligoxanthum , with which this subspecies co-occurs in many parts of its distribution. Reliably distinguished only by shape of the apex of aedeagus, pointed in O. picturatum and O. oligoxanthum , but obtuse in O. capense ( Figs. 39–45 ).

Redescription. Body length 5.02–6.05 mm; width 3.35–4.03 mm. Habitus ( Fig. 27–30 ).

Color: Elytra, sides of pronotum and head yellow. Labrum, antennae and legs flavescent; pattern on elytra, pronotum and head metallic green. Venter, excluding epipleura, dark rufopiceous. Proepipleura and elytral epipleura pale piceous.

Head moderately flat, coarsely punctate laterally and posteriorly, more sparsely punctate around clypeus and on middle of frons, with punctures deep and with sides distinctly margined. Pale pattern on head V-shaped and well-developed. Clypeus with small wrinkles and bisetose, anterior margin formed by two straight lines meeting at an obtuse angle. Labrum slightly bisinuate, with six setae. Subocular ridge well developed. Antennal scape unisetose at apex, antennomeres 3 and 4 with 4–6 setae, antennomeres 5–11 pubescent. HW: 1.59–1.75 mm.

Pronotum (PL: 1.18–1.37 mm, PW: 2.63–2.91 mm.) moderately convex, transverse, base bisinuate, sides slightly rounded, hind angles almost acute. Sides of pronotum with relatively wide border. Pronotum sparsely but coarsely punctate, punctures denser along anterior and posterior margins; metallic green pattern wide, anterior margin more or less yellowish, lateral margins with broad yellow edging. Median line almost obsolete or reduced. Prosternum coarsely and irregularly punctate.

Elytra (EL: 3.18–3.49 mm, EW: 3.35–4.0 mm) moderately convex, ovate, sides hardly widened behind humerus. Elytra with 15 striae, extended from base to apex. Striae deep, punctate, with punctures distinct, well separated, shallower toward apex. Intervals moderately convex and smooth, more convex and narrower laterally.

Venter ( Fig. 35 ). Mesepisternum smooth. Metasternum coarsely and irregularly punctate. Ventrite I punctate along anterior margin; ventrite II with few punctures near coxa and near elytral epipleura; ventrites III–VI smooth. Metacoxae bisetose, without additional punctures.

Aedeagus ( Figs. 39–41 ). Apical lamella slightly downturned, apex relatively wide in comparison with other subspecies. Endophallus is typical for the species: with two groups of medium sized spines. One group is large and situated in distal part tubular formation, extended to basal orifice of median lobe, and covered with scales. The second group of spines is much smaller, located more apically and to the right (anatomically) of the first and extended apically beyond the first group by about half its length ( Fig. 46 ).

Comments. Omophron dominicense was described by Chaudoir, 1868: 58 based on a single specimen received from A. Deyrolle with label “S Domingo” [label error]. The type specimen was studied by S.W. Nichols in 1983 and synonymy with O. capensis was established, but not published. This synonymy is confirmed here.

The population living on the coast, in the Cape Town area, is distinguished by a slightly larger body size and a reduced green pattern on the elytra ( Fig. 27–28 ), and was described by Guérin-Méneville (1842: 24) as Omophron suturale . No differences were found in the structure of the aedeagus. The previously proposed by S.W. Nichols, but unpublished synonymy is confirmed.

Distribution. Botswana, Mozambique, Namibia, Republic of South Africa, Zambia, Zimbabwe (Map. 3).

References: Csiki 1927; Gory 1833; Guérin-Méneville 1842; Péringuey 1896.

Figures

FIGURES 39–45. Aedeagus structure. a—right lateral view; b—apex, dorsal view:: 39—O. (O.) capense s.str. (RSA); 40— O. dominicense Chaudoir, 1868 syn. n. (Type); 41—O. (O.) capense s.str. (f. suturale); 42—O. (O.) capense kmecoi ssp. n. (Holotype); 43—O. (O.) capense congoense stat. n. (Paratype); 44—O. (O.) capense pumilum ssp. n. (Paratype); 45—O. (O.) capense isolatum ssp. n. (Paratype). Scale bar 1 mm.

FIGURES 27–34. Habitus of subspecies of O. (O.) capense. dorsal view: 27—O. (O.) capense s.str. (Type); 28—O. (O.) capense s.str. (f. suturale); 29—O. (O.) capense s.str. (inland form, RSA); 30—O. dominicense Chaudoir, 1868 syn. n. (Type); 31—O. (O.) capense congoense stat. n. (Paratype); 32—O. (O.) capense pumilum ssp. n. (Paratype); 33—O. (O.) capense kmecoi ssp. n. (Holotype); 34—O. (O.) capense isolatum ssp. n. (Paratype). Scale bar 5 mm.

FIGURES 35–38. Habitus of subspecies of O. (O.) capense. ventral view: 35—O. (O.) capense s.str.; 36—O. (O.) capense pumilum ssp. n. (Paratype); 37—O. (O.) capense kmecoi ssp. n. (Paratype); 38—O. (O.) capense isolatum ssp. n. (Paratype). Scale bar 5 mm.

FIGURES 46–51. Endofallus structure of subspecies of O. (O.) capense. dorsal view: 46—O. (O.) capense s.str. (f. suturale); 47—O. (O.) capense kmecoi ssp. n. (Holotype); 48—O. (O.) capense congoense stat. n. (Paratype); 49—O. (O.) capense pumilum ssp. n. (Paratype; Angola: Kangube); 50—O. (O.) capense pumilum ssp. n. (Paratype; Namibia: Caprivi); 51—O. (O.) capense isolatum ssp. n. (Paratype). Scale bar 1 mm.