Syspasis scutellator ( GRAVENHORST , 1829)

Riedel, Matthias, 2021, Revision of the Palaearctic species of Syspasis TOWNES, 1965 (Hymenoptera, Ichneumonidae, Ichneumoninae), Linzer biologische Beiträge 53 (2), pp. 775-810 : 793-794

Syspasis scutellator ( GRAVENHORST, 1829) (figs 11, 22, 31, 44, 58)

Original: Ichneumon scutellator GRAVENHORST, 1829: 175 . L e c t o t y p e: (♀) OE. (Wroclaw), type not studied. syn. Ichneumon scutellator var. iberica SEYRIG, 1927 , synonymized by SELFA (1995: 436). M a t e r i a l e x a m i n e d: 99♀♀ and 48♁♁ from Andorra, Austria, Belgium, Bulgaria,

Czech Republic, Germany, France, Greece *, Italy, Kyrgyzstan *, Poland, Romania, Russia

(Karelia, Altai, Irkutsk, Russia Far East), Switzerland, Sicily, Spain, Tunesia, Turkey.

D e s c r i p t i o n: ♀: Body length 11-13 mm. Flagellum with 35-38 flagellomeres, moderately lanceolate; 1 st flagellomere 3.3-3.8× and 2 nd flagellomere 2.6-2.8× longer than wide, combined length 0.78-0.85× eye length; widest flagellomeres 1.2-1.5× wider than long. Temple parallel behind eye and slightly narrowed apically. OED 1.5-2.0× ocellar diameter. Frons finely punctate or rugose-punctate. Face punctate and granulate. Clypeus large, 1.8× wider than long, apical margins slightly thickened and slightly impressed medially. Gena densely punctate, 1.1× wider than eye. MI 1.0-1.2. Lower mandibular teeth moderately turned inwards (by 45°). Genal carina reaching hypostomal carina close to mandibular base; hypostomal carina slightly elevated. Mesosoma covered with short whitish hairs. Side of pronotum punctate dorsally and striate ventrally. Notaulus impressed frontally. Lateral field of mesoscutum densely punctate and granulate. Mesopleuron densely punctate, partly rugose. Metapleuron punctate, partly rugose, juxtacoxal carina ± distinct. Scutellum wider than long, moderately elevated, densely punctate, without lateral carina. Area basalis with small and sometimes indistinct median tubercle. Area superomedia heart-shaped, as long or slightly longer than wide, rugose; anterior transverse carina sometimes indistinct, in or caudal to its middle. Area externa punctate and granulate. Area petiolaris rugose or transversely striate, with lateral carina. Hind coxa densely punctate and granulate, with small, often diffuse apico-ventral scopa (c.0.25× coxal length). Hind femur densely punctate, (4.0-) 4.3-4.5× longer than wide. Areolet pentagonal, frontal distance between veins 2rs-m and 3rs-m 1-2× their diameter; vein 2m-cu reaching its middle. Vein 1cu-a interstitial or slightly postfurcal. Metasoma oxypygous. Lateral field of postpetiolus rugose-punctate, 0.6-0.8× as wide as median field, the latter with fine longitudinal striae or rugae. Gastrocoelus distinct, with longitudinal ridges. Thyridium transverse or slightly oblique, 1.5-3.0× wider than the interval between thyridia. 2 nd tergite 0.85-1.0× as long as wide. 2 nd and 3 rd tergites densely rugose-punctate, ± dull. 4 th and 5 th tergites with superficial punctures, 5 th tergite often almost smooth. Ovipositor sheath slightly widened apically and distinctly extending metasomal apex (by 0.8-1.0× length of 7 th tergite).

Color: black. Basal flagellomeres often pale apically or ± reddish. Stripes or rings on flagellomeres 7/8-12 (sometimes reduced), sometimes side of clypeus, frontal orbit, small spot on vertex, rarely collar, sometimes upper margin or hind edge of pronotum, sometimes spot on subtegular ridge, side of scutellum, rarely postscutellum, often fore femur apically and fore tibia frontally ivory. Coxae and trochanters black, sometimes fore and mid coxae ± reddish. Legs otherwise red; hind tarsus ± infuscate. Tergites black, sometimes postpetiolus, 2 nd and 3 rd tergites and base of 4 th tergite dark reddish or with reddish apical bands. Wings slightly infuscate; pterostigma yellowish or ochreous.

♁: Body length 12-13 mm. Flagellum with 35-38 flagellomeres; 1 st flagellomere 2.5× and 2 nd flagellomere 2.1× longer than wide, combined length 0.65× eye length. Tyloids on flagellomeres 7/9-15/17, linear, maximally as long as their flagellomeres. OED 1.0- 1.5× ocellar diameter. Frons transversely rugose-punctate. Gena 1.1-1.3× as wide as eye. MI 0.6-0.7. Mesosoma with short pale hairs. Scutellum wider than long, moderately elevated, densely punctate, with lateral carina in basal 0.2-0.5. Coxae densely punctate and granulate, with dense pilosity. Hind femur 3.9-4.1× longer than wide. Fore metatarsus without apical tooth. Lateral field of postpetiolus granulate, 0.5-0.7× as wide as median field. Thyridium slightly oblique, 1.5-2.5× wider than the interval between thyridia. 2 nd tergite 1.0-1.1× longer than wide. 2 nd to 4 th /5 th tergites superficially rugose-punctate, 2 nd tergite basally striate.

Color: black. Flagellum reddish or ochreous ventrally. Usually face and clypeus (sometimes except black median stripes), mandibular base, palps, frontal orbit, stripe on vertex, often spot on outer orbit, usually scape ventrally, collar and upper margin of pronotum, subtegular ridge, side of scutellum, often postscutellum, usually fore and mid coxae mainly, and sometimes apical spot on postpetiolus yellow. 2 nd to 4 th tergites often diffusely dark reddish. Coxae black, trochanters often ± reddish. Legs otherwise red; fore and mid tibiae yellowish frontally, hind tibia often infuscate apically; hind tarsus blackish. Pterostigma yellow or brownish.

Hosts: Euproctis chrysorrhoea (LINNAEUS, 1758) (Lep., Erebidae ), Abraxas grossulariata (LINNAEUS, 1758) (Lep., Geometridae ), Panolis flammea (DENIS & SCHIFFERMÜLLER, 1775) (Lep., Noctuidae ) ( RASNITSYN & SIYTAN 1981).